Articles | Volume 22, issue 6
https://doi.org/10.5194/cp-22-1125-2026
© Author(s) 2026. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
https://doi.org/10.5194/cp-22-1125-2026
© Author(s) 2026. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
Unravelling the tree cover dynamics over the last 20 000 years on the Northern Hemisphere
Anne Dallmeyer
CORRESPONDING AUTHOR
Max-Planck-Institute for Meteorology, 20146 Hamburg, Germany
Nils Weitzel
Research Center Trustworthy Data Science and Security of the University Alliance Ruhr, Dortmund, Germany
Department of Statistics, TU Dortmund University, Dortmund, Germany
School of Geographical Sciences, University of Bristol, Bristol, UK
Laura Schild
Helmholtz Centre for Polar and Marine Research, Research Unit Potsdam, Alfred Wegener Institute (AWI), Potsdam, Germany
Ulrike Herzschuh
Helmholtz Centre for Polar and Marine Research, Research Unit Potsdam, Alfred Wegener Institute (AWI), Potsdam, Germany
Institute of Environmental Sciences and Geography, University of Potsdam, Karl-Liebknecht-Straße 24–25, Potsdam, Germany
Institute of Biochemistry and Biology, University of Potsdam, Karl-Liebknecht-Straße 24–25, Potsdam, Germany
Thomas Kleinen
Max-Planck-Institute for Meteorology, 20146 Hamburg, Germany
Martin Claussen
Max-Planck-Institute for Meteorology, 20146 Hamburg, Germany
Meteorological Institute, Universität Hamburg, 20146 Hamburg, Germany
Related authors
Martin Claussen, Anne Dallmeyer, Frank Darius, Philipp Hoelzmann, and Michèle Dinies
EGUsphere, https://doi.org/10.5194/egusphere-2026-2587, https://doi.org/10.5194/egusphere-2026-2587, 2026
This preprint is open for discussion and under review for Climate of the Past (CP).
Short summary
Short summary
The abrupt termination of humid conditions and vegetation changes in the Sahara during the last 8000 years is an important issue as it presumably affected the time frame for human adaption. Our analysis suggests that there was no “collapse” of the green Sahara. Gradual changes in some proxy records and abrupt shifts in others are not necessarily contradictory and a gradual expansion of the Sahara may have been accompanied by abrupt changes within the ecosystems.
Chenzhi Li, Anne Dallmeyer, Jian Ni, Manuel Chevalier, Matteo Willeit, Andrei A. Andreev, Xianyong Cao, Laura Schild, Birgit Heim, Mareike Wieczorek, and Ulrike Herzschuh
Clim. Past, 21, 1001–1024, https://doi.org/10.5194/cp-21-1001-2025, https://doi.org/10.5194/cp-21-1001-2025, 2025
Short summary
Short summary
We present global megabiome dynamics and distributions derived from pollen-based reconstructions over the last 21 000 years, which are suitable for the evaluation of Earth-system-model-based paleo-megabiome simulations. We identified strong deviations between pollen- and model-derived megabiome distributions in the circum-Arctic and Tibetan Plateau areas during the Last Glacial Maximum and early deglaciation and in northern Africa and the Mediterranean region during the Holocene.
Anne Dallmeyer, Anneli Poska, Laurent Marquer, Andrea Seim, and Marie-José Gaillard
Clim. Past, 19, 1531–1557, https://doi.org/10.5194/cp-19-1531-2023, https://doi.org/10.5194/cp-19-1531-2023, 2023
Short summary
Short summary
We compare past tree cover changes in Europe during the last 8000 years simulated with two dynamic global vegetation models and inferred from pollen data. The major model–data mismatch is related to the much earlier onset of anthropogenic deforestation in the data compared to the prescribed land use in the models. We show that land use, and not climate, is the main driver of the Holocene forest decline. The model–data agreement depends on the model tuning, challenging model–data comparisons.
Ulrike Herzschuh, Thomas Böhmer, Manuel Chevalier, Raphaël Hébert, Anne Dallmeyer, Chenzhi Li, Xianyong Cao, Odile Peyron, Larisa Nazarova, Elena Y. Novenko, Jungjae Park, Natalia A. Rudaya, Frank Schlütz, Lyudmila S. Shumilovskikh, Pavel E. Tarasov, Yongbo Wang, Ruilin Wen, Qinghai Xu, and Zhuo Zheng
Clim. Past, 19, 1481–1506, https://doi.org/10.5194/cp-19-1481-2023, https://doi.org/10.5194/cp-19-1481-2023, 2023
Short summary
Short summary
A mismatch between model- and proxy-based Holocene climate change may partially originate from the poor spatial coverage of climate reconstructions. Here we investigate quantitative reconstructions of mean annual temperature and annual precipitation from 1908 pollen records in the Northern Hemisphere. Trends show strong latitudinal patterns and differ between (sub-)continents. Our work contributes to a better understanding of the global mean.
Ulrike Herzschuh, Thomas Böhmer, Chenzhi Li, Manuel Chevalier, Raphaël Hébert, Anne Dallmeyer, Xianyong Cao, Nancy H. Bigelow, Larisa Nazarova, Elena Y. Novenko, Jungjae Park, Odile Peyron, Natalia A. Rudaya, Frank Schlütz, Lyudmila S. Shumilovskikh, Pavel E. Tarasov, Yongbo Wang, Ruilin Wen, Qinghai Xu, and Zhuo Zheng
Earth Syst. Sci. Data, 15, 2235–2258, https://doi.org/10.5194/essd-15-2235-2023, https://doi.org/10.5194/essd-15-2235-2023, 2023
Short summary
Short summary
Climate reconstruction from proxy data can help evaluate climate models. We present pollen-based reconstructions of mean July temperature, mean annual temperature, and annual precipitation from 2594 pollen records from the Northern Hemisphere, using three reconstruction methods (WA-PLS, WA-PLS_tailored, and MAT). Since no global or hemispheric synthesis of quantitative precipitation changes are available for the Holocene so far, this dataset will be of great value to the geoscientific community.
Manuel Chevalier, Anne Dallmeyer, Nils Weitzel, Chenzhi Li, Jean-Philippe Baudouin, Ulrike Herzschuh, Xianyong Cao, and Andreas Hense
Clim. Past, 19, 1043–1060, https://doi.org/10.5194/cp-19-1043-2023, https://doi.org/10.5194/cp-19-1043-2023, 2023
Short summary
Short summary
Data–data and data–model vegetation comparisons are commonly based on comparing single vegetation estimates. While this approach generates good results on average, reducing pollen assemblages to single single plant functional type (PFT) or biome estimates can oversimplify the vegetation signal. We propose using a multivariate metric, the Earth mover's distance (EMD), to include more details about the vegetation structure when performing such comparisons.
Anne Dallmeyer, Martin Claussen, Stephan J. Lorenz, Michael Sigl, Matthew Toohey, and Ulrike Herzschuh
Clim. Past, 17, 2481–2513, https://doi.org/10.5194/cp-17-2481-2021, https://doi.org/10.5194/cp-17-2481-2021, 2021
Short summary
Short summary
Using the comprehensive Earth system model, MPI-ESM1.2, we explore the global Holocene vegetation changes and interpret them in terms of the Holocene climate change. The model results reveal that most of the Holocene vegetation transitions seen outside the high northern latitudes can be attributed to modifications in the intensity of the global summer monsoons.
Mengze Li, Robert B. Jackson, Marielle Saunois, Philippe Ciais, Ben Poulter, Josep G. Canadell, Prabir K. Patra, Hanqin Tian, Zhen Zhang, Etienne Fluet-Chouinard, Zutao Ouyang, Ting Zhang, David J. Beerling, Dmitry A. Belikov, Philippe Bousquet, Danilo Custodio, Naveen Chandra, Xinyu Dou, Nicola Gedney, Peter O. Hopcroft, Alison M. Hoyt, Kazuhito Ichii, Akihito Ito, Atul K. Jain, Katherine Jensen, Fortunat Joos, Thomas Kleinen, Masayuki Kondo, Fa Li, Tingting Li, Xiangyu Liu, Shamil Maksyutov, Avni Malhotra, Adrien Martinez, Kyle McDonald, Joe R. Melton, Jurek Müller, Yosuke Niwa, Shufen Pan, Shushi Peng, Changhui Peng, Zhangcai Qin, Peter Raymond, William Riley, Arjo Segers, Rona L. Thompson, Aki Tsuruta, Yi Xi, Kunxiaojia Yuan, Wenxin Zhang, Bo Zheng, Qing Zhu, Qiuan Zhu, and Qianlai Zhuang
Earth Syst. Sci. Data, 18, 3507–3524, https://doi.org/10.5194/essd-18-3507-2026, https://doi.org/10.5194/essd-18-3507-2026, 2026
Short summary
Short summary
We proposed a framework that combines machine-learning and climate data to predict global natural vegetated wetland methane emissions for 2000–2025. We found that although total global emissions remained stable in the post-2020s, Northern Hemisphere emissions surged whilst tropical emissions fell. This approach allows us to rapidly monitor emissions and provides early warnings for climate impacts.
Martin Claussen, Anne Dallmeyer, Frank Darius, Philipp Hoelzmann, and Michèle Dinies
EGUsphere, https://doi.org/10.5194/egusphere-2026-2587, https://doi.org/10.5194/egusphere-2026-2587, 2026
This preprint is open for discussion and under review for Climate of the Past (CP).
Short summary
Short summary
The abrupt termination of humid conditions and vegetation changes in the Sahara during the last 8000 years is an important issue as it presumably affected the time frame for human adaption. Our analysis suggests that there was no “collapse” of the green Sahara. Gradual changes in some proxy records and abrupt shifts in others are not necessarily contradictory and a gradual expansion of the Sahara may have been accompanied by abrupt changes within the ecosystems.
Moritz Adam, Elisa Ziegler, Björn Gonzalez, Nils Weitzel, and Kira Rehfeld
EGUsphere, https://doi.org/10.5194/egusphere-2026-626, https://doi.org/10.5194/egusphere-2026-626, 2026
This preprint is open for discussion and under review for Earth System Dynamics (ESD).
Short summary
Short summary
As plants take up Carbon Dioxide from the atmosphere, they buffer climate change. They are vulnerable to extreme weather and climate. When temperature and moisture extremes happen simultaneously, they strongly affect how plants grow. We use complex computer models to understand impacts of extremes on plants. We compared the models to observations. The models do well globally and okay in many regions. Locally, they need improvement because modeling how water moves through the soil is difficult.
Lin Yu, Thomas Kleinen, Min Jung Kwon, Christian Knoblauch, and Christian Beer
EGUsphere, https://doi.org/10.5194/egusphere-2025-4648, https://doi.org/10.5194/egusphere-2025-4648, 2025
Short summary
Short summary
We studied how adding biochar to soils might affect future climate. Using computer simulations, we found that while global averages of temperature and rainfall change little, extreme events respond more clearly. Heat waves and heavy rain are reduced in many regions, though drought risks rise in some dry areas. These results suggest that biochar could help moderate harmful climate extremes, especially on land, but with region-specific effects.
Ingolf Kühn, Christian Hecht, Ulrike Herzschuh, and Dirk Scherler
Web Ecol., 25, 157–168, https://doi.org/10.5194/we-25-157-2025, https://doi.org/10.5194/we-25-157-2025, 2025
Short summary
Short summary
Since 1850, glaciers have retreated in the Alps, providing ground for vegetation succession. Such systems were studied intensively in other parts of the Alps, but excluding the Northern Limestone Alps. Hence, we initiated a long-term research programme, which we introduce here. Initial findings show an increase in plant species richness and cover with age since deglaciation. This is, however, by far slower than observed elsewhere in the Alps, likely due to the specific geology and geomorphology.
Trevor Martin Sloughter, Zebedee Nicholls, Gang Tang, Thomas Kleinen, Zhen Zhang, and Joeri Rogelj
EGUsphere, https://doi.org/10.5194/egusphere-2025-3873, https://doi.org/10.5194/egusphere-2025-3873, 2025
Short summary
Short summary
High resolution models of the earth system exhibit some disagreement and uncertainty on future methane emissions from natural sources, in particular wetlands, with some studies predicting wetlands alone could be very significant sources over the 21st century. Modelling these emissions as a response to global temperature is one option for simple models to approximate the climate impact of wetlands. The effect is a small increase in overall temperatures and a widening of the uncertainty range.
Izabella A. Baisheva, Birgit Heim, Jorge García Molinos, Amelie Stieg, Hanno Meyer, Ramesh Glückler, Kathleen R. Stoof-Leichsenring, Antje Eulenburg, Pier Paul Overduin, Evgenii S. Zakharov, Aital V. Egorov, Paraskovya V. Davydova, Lena A. Ushnitskaya, Sardana N. Levina, Ruslan M. Gorodnichev, Robert Jackisch, Antonie Haas, Stefan Kruse, Luidmila A. Pestryakova, and Ulrike Herzschuh
EGUsphere, https://doi.org/10.5194/egusphere-2025-4114, https://doi.org/10.5194/egusphere-2025-4114, 2025
Short summary
Short summary
Our study provides a new comprehensive assessment of the limnological state of 66 lakes in the Central Yakutian alaas landscapes and the Verkhoyansk mountain range. Our analyses suggest that specific lake-type properties within the thermokarst lake sequence seem to drive inorganic, organic, and isotopic lake hydrochemistry. Future warming may lead to less diversification within lake macrophyte assemblages in old alaas lakes.
Stiig Wilkenskjeld, Thomas Kleinen, Tobias Stacke, and Victor Brovkin
EGUsphere, https://doi.org/10.5194/egusphere-2025-3601, https://doi.org/10.5194/egusphere-2025-3601, 2025
Short summary
Short summary
Methane is the second most important greenhouse gas with high potential for short term reductions of human induced global warming. We model methane emissions from the most important and most uncertain natural source: wetlands. We investigate how a number of assumptions, including human impact on natural wetlands, influences the wetlands and their methane emissions. Of the tested influences we find the most important to be how humans are altering the soil surface.
Lutz Schirrmeister, Margret C. Fuchs, Thomas Opel, Andrei Andreev, Frank Kienast, Andrea Schneider, Larisa Nazarova, Larisa Frolova, Svetlana Kuzmina, Tatiana Kuznetsova, Vladimir Tumskoy, Heidrun Matthes, Gerrit Lohmann, Guido Grosse, Viktor Kunitsky, Hanno Meyer, Heike H. Zimmermann, Ulrike Herzschuh, Thomas Böhmer, Stuart Umbo, Sevi Modestou, Sebastian F. M. Breitenbach, Anfisa Pismeniuk, Georg Schwamborn, Stephanie Kusch, and Sebastian Wetterich
Clim. Past, 21, 1143–1184, https://doi.org/10.5194/cp-21-1143-2025, https://doi.org/10.5194/cp-21-1143-2025, 2025
Short summary
Short summary
Geochronological, cryolithological, paleoecological, and modeling data reconstruct the Last Interglacial (LIG) climate around the New Siberian Islands and reveal significantly warmer conditions compared to today. The critical challenges in predicting future ecosystem responses lie in the fact that the land–ocean distribution during the LIG was markedly different from today, affecting the degree of continentality, which played a major role in modulating climate and ecosystem dynamics.
Ulrike Herzschuh, Thomas Böhmer, Weihan Jia, and Simeon Lisovski
EGUsphere, https://doi.org/10.5194/egusphere-2025-2678, https://doi.org/10.5194/egusphere-2025-2678, 2025
Short summary
Short summary
We introduce a new climate proxy based on plant DNA preserved in lake sediments. Validated with a large surface sample dataset and applied to a sediment record, this method provides more accurate and robust reconstructions of past climate change than traditional vegetation proxies like pollen, likely due to a higher taxonomic resolution and more localized signal.
Chenzhi Li, Anne Dallmeyer, Jian Ni, Manuel Chevalier, Matteo Willeit, Andrei A. Andreev, Xianyong Cao, Laura Schild, Birgit Heim, Mareike Wieczorek, and Ulrike Herzschuh
Clim. Past, 21, 1001–1024, https://doi.org/10.5194/cp-21-1001-2025, https://doi.org/10.5194/cp-21-1001-2025, 2025
Short summary
Short summary
We present global megabiome dynamics and distributions derived from pollen-based reconstructions over the last 21 000 years, which are suitable for the evaluation of Earth-system-model-based paleo-megabiome simulations. We identified strong deviations between pollen- and model-derived megabiome distributions in the circum-Arctic and Tibetan Plateau areas during the Last Glacial Maximum and early deglaciation and in northern Africa and the Mediterranean region during the Holocene.
Yana Savytska, Viktor Smolii, and Nils Weitzel
Earth Syst. Dynam., 16, 721–727, https://doi.org/10.5194/esd-16-721-2025, https://doi.org/10.5194/esd-16-721-2025, 2025
Short summary
Short summary
In recent decades, we have witnessed abnormally hot summers and frequent weather extremes globally. These are clear signs of global warming and climate change. A constant increase in atmospheric carbon dioxide (CO2) is a major driver of these changes. We propose an algorithm for near-real-time detection of terrestrial areas with CO2 sources and sinks. This algorithm could aid in developing new methods of natural CO2 reduction and exploring ecosystem responses to disturbances.
Amelie Stieg, Boris K. Biskaborn, Ulrike Herzschuh, Andreas Marent, Jens Strauss, Dorothee Wilhelms-Dick, Luidmila A. Pestryakova, and Hanno Meyer
Biogeosciences, 22, 2327–2350, https://doi.org/10.5194/bg-22-2327-2025, https://doi.org/10.5194/bg-22-2327-2025, 2025
Short summary
Short summary
Globally, lake ecosystems have undergone significant shifts since the 1950s due to human activities. This study presents a unique ~220-year sediment record from a remote Siberian boreal lake, providing a multiproxy perspective on climate warming and anthropogenic air pollution. Analyses of diatom assemblages, diatom silicon isotopes, and carbon and nitrogen sediment proxies reveal complex biogeochemical interactions, highlighting anthropogenic influences even on remote water resources.
Laura Schild, Peter Ewald, Chenzhi Li, Raphaël Hébert, Thomas Laepple, and Ulrike Herzschuh
Earth Syst. Sci. Data, 17, 2007–2033, https://doi.org/10.5194/essd-17-2007-2025, https://doi.org/10.5194/essd-17-2007-2025, 2025
Short summary
Short summary
This study reconstructed vegetation and tree cover in the Northern Hemisphere from a harmonized dataset of pollen counts from sediment and peat cores for the past 14 000 years. A model was applied to correct for differences in pollen production between different plants, and modern remote-sensing forest cover was used to validate the reconstructed tree cover. Accurate data on past vegetation are invaluable for the investigation of vegetation–climate dynamics and the validation of vegetation models.
Marielle Saunois, Adrien Martinez, Benjamin Poulter, Zhen Zhang, Peter A. Raymond, Pierre Regnier, Josep G. Canadell, Robert B. Jackson, Prabir K. Patra, Philippe Bousquet, Philippe Ciais, Edward J. Dlugokencky, Xin Lan, George H. Allen, David Bastviken, David J. Beerling, Dmitry A. Belikov, Donald R. Blake, Simona Castaldi, Monica Crippa, Bridget R. Deemer, Fraser Dennison, Giuseppe Etiope, Nicola Gedney, Lena Höglund-Isaksson, Meredith A. Holgerson, Peter O. Hopcroft, Gustaf Hugelius, Akihiko Ito, Atul K. Jain, Rajesh Janardanan, Matthew S. Johnson, Thomas Kleinen, Paul B. Krummel, Ronny Lauerwald, Tingting Li, Xiangyu Liu, Kyle C. McDonald, Joe R. Melton, Jens Mühle, Jurek Müller, Fabiola Murguia-Flores, Yosuke Niwa, Sergio Noce, Shufen Pan, Robert J. Parker, Changhui Peng, Michel Ramonet, William J. Riley, Gerard Rocher-Ros, Judith A. Rosentreter, Motoki Sasakawa, Arjo Segers, Steven J. Smith, Emily H. Stanley, Joël Thanwerdas, Hanqin Tian, Aki Tsuruta, Francesco N. Tubiello, Thomas S. Weber, Guido R. van der Werf, Douglas E. J. Worthy, Yi Xi, Yukio Yoshida, Wenxin Zhang, Bo Zheng, Qing Zhu, Qiuan Zhu, and Qianlai Zhuang
Earth Syst. Sci. Data, 17, 1873–1958, https://doi.org/10.5194/essd-17-1873-2025, https://doi.org/10.5194/essd-17-1873-2025, 2025
Short summary
Short summary
Methane (CH4) is the second most important human-influenced greenhouse gas in terms of climate forcing after carbon dioxide (CO2). A consortium of multi-disciplinary scientists synthesise and update the budget of the sources and sinks of CH4. This edition benefits from important progress in estimating emissions from lakes and ponds, reservoirs, and streams and rivers. For the 2010s decade, global CH4 emissions are estimated at 575 Tg CH4 yr-1, including ~65 % from anthropogenic sources.
Simeon Lisovski, Alexandra Runge, Iuliia Shevtsova, Nele Landgraf, Anne Morgenstern, Ronald Reagan Okoth, Matthias Fuchs, Nikolay Lashchinskiy, Carl Stadie, Alison Beamish, Ulrike Herzschuh, Guido Grosse, and Birgit Heim
Earth Syst. Sci. Data, 17, 1707–1730, https://doi.org/10.5194/essd-17-1707-2025, https://doi.org/10.5194/essd-17-1707-2025, 2025
Short summary
Short summary
The Lena Delta is the largest river delta in the Arctic and represents a biodiversity hotspot. Here, we describe multiple field datasets and a detailed habitat classification map for the Lena Delta. We present context and methods of these openly available datasets and show how they can improve our understanding of the rapidly changing Arctic tundra system.
Mateo Duque-Villegas, Martin Claussen, Thomas Kleinen, Jürgen Bader, and Christian H. Reick
Clim. Past, 21, 773–794, https://doi.org/10.5194/cp-21-773-2025, https://doi.org/10.5194/cp-21-773-2025, 2025
Short summary
Short summary
We simulate the last glacial cycle with a comprehensive model of the Earth system and investigate vegetation cover change in northern Africa during the last four African Humid Periods (AHPs). We find a common pattern of vegetation change and relate it with climatic factors in order to discuss how vegetation might have evolved during even older AHPs. This scaling relationship we find according to past AHPs fails to account for projected changes in northern Africa under strong greenhouse gas warming.
Elisa Ziegler, Nils Weitzel, Jean-Philippe Baudouin, Marie-Luise Kapsch, Uwe Mikolajewicz, Lauren Gregoire, Ruza Ivanovic, Paul J. Valdes, Christian Wirths, and Kira Rehfeld
Clim. Past, 21, 627–659, https://doi.org/10.5194/cp-21-627-2025, https://doi.org/10.5194/cp-21-627-2025, 2025
Short summary
Short summary
During the Last Deglaciation, global surface temperature rose by about 4–7 °C over several millennia. We show that changes in year-to-year up to century-to-century fluctuations of temperature and precipitation during the Deglaciation were mostly larger than during either the preceding or succeeding more stable periods in 15 climate model simulations. The analysis demonstrates how ice sheets, meltwater, and volcanism influence simulated variability to inform future simulation protocols.
Jean-Philippe Baudouin, Nils Weitzel, Maximilian May, Lukas Jonkers, Andrew M. Dolman, and Kira Rehfeld
Clim. Past, 21, 381–403, https://doi.org/10.5194/cp-21-381-2025, https://doi.org/10.5194/cp-21-381-2025, 2025
Short summary
Short summary
Earth's past temperature reconstructions are critical for understanding climate change. We test the ability of these reconstructions using climate simulations. Uncertainties, mainly from past temperature measurement methods and age determination, impact reconstructions over time. While more data enhance accuracy for long-term trends, high-quality data are more important for short-term precision. Our study lays the groundwork for better reconstructions and suggests avenues for improvement.
Tuula Aalto, Aki Tsuruta, Jarmo Mäkelä, Jurek Müller, Maria Tenkanen, Eleanor Burke, Sarah Chadburn, Yao Gao, Vilma Mannisenaho, Thomas Kleinen, Hanna Lee, Antti Leppänen, Tiina Markkanen, Stefano Materia, Paul A. Miller, Daniele Peano, Olli Peltola, Benjamin Poulter, Maarit Raivonen, Marielle Saunois, David Wårlind, and Sönke Zaehle
Biogeosciences, 22, 323–340, https://doi.org/10.5194/bg-22-323-2025, https://doi.org/10.5194/bg-22-323-2025, 2025
Short summary
Short summary
Wetland methane responses to temperature and precipitation were studied in a boreal wetland-rich region in northern Europe using ecosystem models, atmospheric inversions, and upscaled flux observations. The ecosystem models differed in their responses to temperature and precipitation and in their seasonality. However, multi-model means, inversions, and upscaled fluxes had similar seasonality, and they suggested co-limitation by temperature and precipitation.
Zhen Zhang, Benjamin Poulter, Joe R. Melton, William J. Riley, George H. Allen, David J. Beerling, Philippe Bousquet, Josep G. Canadell, Etienne Fluet-Chouinard, Philippe Ciais, Nicola Gedney, Peter O. Hopcroft, Akihiko Ito, Robert B. Jackson, Atul K. Jain, Katherine Jensen, Fortunat Joos, Thomas Kleinen, Sara H. Knox, Tingting Li, Xin Li, Xiangyu Liu, Kyle McDonald, Gavin McNicol, Paul A. Miller, Jurek Müller, Prabir K. Patra, Changhui Peng, Shushi Peng, Zhangcai Qin, Ryan M. Riggs, Marielle Saunois, Qing Sun, Hanqin Tian, Xiaoming Xu, Yuanzhi Yao, Yi Xi, Wenxin Zhang, Qing Zhu, Qiuan Zhu, and Qianlai Zhuang
Biogeosciences, 22, 305–321, https://doi.org/10.5194/bg-22-305-2025, https://doi.org/10.5194/bg-22-305-2025, 2025
Short summary
Short summary
This study assesses global methane emissions from wetlands between 2000 and 2020 using multiple models. We found that wetland emissions increased by 6–7 Tg CH4 yr-1 in the 2010s compared to the 2000s. Rising temperatures primarily drove this increase, while changes in precipitation and CO2 levels also played roles. Our findings highlight the importance of wetlands in the global methane budget and the need for continuous monitoring to understand their impact on climate change.
Pin-Hsin Hu, Christian H. Reick, Reiner Schnur, Axel Kleidon, and Martin Claussen
Geosci. Model Dev. Discuss., https://doi.org/10.5194/gmd-2024-111, https://doi.org/10.5194/gmd-2024-111, 2024
Revised manuscript accepted for GMD
Short summary
Short summary
We introduce the new plant functional diversity model JeDi-BACH, a novel tool that integrates the Jena Diversity Model (JeDi) within the land component of the ICON Earth System Model. JeDi-BACH captures a richer set of plant trait variations based on environmental filtering and functional tradeoffs without a priori knowledge of the vegetation types. JeDi-BACH represents a significant advancement in modeling the complex interactions between plant functional diversity and climate.
Nora Farina Specht, Martin Claussen, and Thomas Kleinen
Clim. Past, 20, 1595–1613, https://doi.org/10.5194/cp-20-1595-2024, https://doi.org/10.5194/cp-20-1595-2024, 2024
Short summary
Short summary
We close the terrestrial water cycle across the Sahara and Sahel by integrating a new endorheic-lake model into a climate model. A factor analysis of mid-Holocene simulations shows that both dynamic lakes and dynamic vegetation individually contribute to a precipitation increase over northern Africa that is collectively greater than that caused by the interaction between lake and vegetation dynamics. Thus, the lake–vegetation interaction causes a relative drying response across the entire Sahel.
Amelie Stieg, Boris K. Biskaborn, Ulrike Herzschuh, Jens Strauss, Luidmila Pestryakova, and Hanno Meyer
Clim. Past, 20, 909–933, https://doi.org/10.5194/cp-20-909-2024, https://doi.org/10.5194/cp-20-909-2024, 2024
Short summary
Short summary
Siberia is impacted by recent climate warming and experiences extreme hydroclimate events. We present a 220-year-long sub-decadal stable oxygen isotope record of diatoms from Lake Khamra. Our analysis identifies winter precipitation as the key process impacting the isotope variability. Two possible hydroclimatic anomalies were found to coincide with significant changes in lake internal conditions and increased wildfire activity in the region.
Nils Weitzel, Heather Andres, Jean-Philippe Baudouin, Marie-Luise Kapsch, Uwe Mikolajewicz, Lukas Jonkers, Oliver Bothe, Elisa Ziegler, Thomas Kleinen, André Paul, and Kira Rehfeld
Clim. Past, 20, 865–890, https://doi.org/10.5194/cp-20-865-2024, https://doi.org/10.5194/cp-20-865-2024, 2024
Short summary
Short summary
The ability of climate models to faithfully reproduce past warming episodes is a valuable test considering potentially large future warming. We develop a new method to compare simulations of the last deglaciation with temperature reconstructions. We find that reconstructions differ more between regions than simulations, potentially due to deficiencies in the simulation design, models, or reconstructions. Our work is a promising step towards benchmarking simulations of past climate transitions.
Philip Meister, Anne Alexandre, Hannah Bailey, Philip Barker, Boris K. Biskaborn, Ellie Broadman, Rosine Cartier, Bernhard Chapligin, Martine Couapel, Jonathan R. Dean, Bernhard Diekmann, Poppy Harding, Andrew C. G. Henderson, Armand Hernandez, Ulrike Herzschuh, Svetlana S. Kostrova, Jack Lacey, Melanie J. Leng, Andreas Lücke, Anson W. Mackay, Eniko Katalin Magyari, Biljana Narancic, Cécile Porchier, Gunhild Rosqvist, Aldo Shemesh, Corinne Sonzogni, George E. A. Swann, Florence Sylvestre, and Hanno Meyer
Clim. Past, 20, 363–392, https://doi.org/10.5194/cp-20-363-2024, https://doi.org/10.5194/cp-20-363-2024, 2024
Short summary
Short summary
This paper presents the first comprehensive compilation of diatom oxygen isotope records in lake sediments (δ18OBSi), supported by lake basin parameters. We infer the spatial and temporal coverage of δ18OBSi records and discuss common hemispheric trends on centennial and millennial timescales. Key results are common patterns for hydrologically open lakes in Northern Hemisphere extratropical regions during the Holocene corresponding to known climatic epochs, i.e. the Holocene Thermal Maximum.
Nico Wunderling, Anna S. von der Heydt, Yevgeny Aksenov, Stephen Barker, Robbin Bastiaansen, Victor Brovkin, Maura Brunetti, Victor Couplet, Thomas Kleinen, Caroline H. Lear, Johannes Lohmann, Rosa Maria Roman-Cuesta, Sacha Sinet, Didier Swingedouw, Ricarda Winkelmann, Pallavi Anand, Jonathan Barichivich, Sebastian Bathiany, Mara Baudena, John T. Bruun, Cristiano M. Chiessi, Helen K. Coxall, David Docquier, Jonathan F. Donges, Swinda K. J. Falkena, Ann Kristin Klose, David Obura, Juan Rocha, Stefanie Rynders, Norman Julius Steinert, and Matteo Willeit
Earth Syst. Dynam., 15, 41–74, https://doi.org/10.5194/esd-15-41-2024, https://doi.org/10.5194/esd-15-41-2024, 2024
Short summary
Short summary
This paper maps out the state-of-the-art literature on interactions between tipping elements relevant for current global warming pathways. We find indications that many of the interactions between tipping elements are destabilizing. This means that tipping cascades cannot be ruled out on centennial to millennial timescales at global warming levels between 1.5 and 2.0 °C or on shorter timescales if global warming surpasses 2.0 °C.
Anne Dallmeyer, Anneli Poska, Laurent Marquer, Andrea Seim, and Marie-José Gaillard
Clim. Past, 19, 1531–1557, https://doi.org/10.5194/cp-19-1531-2023, https://doi.org/10.5194/cp-19-1531-2023, 2023
Short summary
Short summary
We compare past tree cover changes in Europe during the last 8000 years simulated with two dynamic global vegetation models and inferred from pollen data. The major model–data mismatch is related to the much earlier onset of anthropogenic deforestation in the data compared to the prescribed land use in the models. We show that land use, and not climate, is the main driver of the Holocene forest decline. The model–data agreement depends on the model tuning, challenging model–data comparisons.
Ulrike Herzschuh, Thomas Böhmer, Manuel Chevalier, Raphaël Hébert, Anne Dallmeyer, Chenzhi Li, Xianyong Cao, Odile Peyron, Larisa Nazarova, Elena Y. Novenko, Jungjae Park, Natalia A. Rudaya, Frank Schlütz, Lyudmila S. Shumilovskikh, Pavel E. Tarasov, Yongbo Wang, Ruilin Wen, Qinghai Xu, and Zhuo Zheng
Clim. Past, 19, 1481–1506, https://doi.org/10.5194/cp-19-1481-2023, https://doi.org/10.5194/cp-19-1481-2023, 2023
Short summary
Short summary
A mismatch between model- and proxy-based Holocene climate change may partially originate from the poor spatial coverage of climate reconstructions. Here we investigate quantitative reconstructions of mean annual temperature and annual precipitation from 1908 pollen records in the Northern Hemisphere. Trends show strong latitudinal patterns and differ between (sub-)continents. Our work contributes to a better understanding of the global mean.
Zoé Rehder, Thomas Kleinen, Lars Kutzbach, Victor Stepanenko, Moritz Langer, and Victor Brovkin
Biogeosciences, 20, 2837–2855, https://doi.org/10.5194/bg-20-2837-2023, https://doi.org/10.5194/bg-20-2837-2023, 2023
Short summary
Short summary
We use a new model to investigate how methane emissions from Arctic ponds change with warming. We find that emissions increase substantially. Under annual temperatures 5 °C above present temperatures, pond methane emissions are more than 3 times higher than now. Most of this increase is caused by an increase in plant productivity as plants provide the substrate microbes used to produce methane. We conclude that vegetation changes need to be included in predictions of pond methane emissions.
Ulrike Herzschuh, Thomas Böhmer, Chenzhi Li, Manuel Chevalier, Raphaël Hébert, Anne Dallmeyer, Xianyong Cao, Nancy H. Bigelow, Larisa Nazarova, Elena Y. Novenko, Jungjae Park, Odile Peyron, Natalia A. Rudaya, Frank Schlütz, Lyudmila S. Shumilovskikh, Pavel E. Tarasov, Yongbo Wang, Ruilin Wen, Qinghai Xu, and Zhuo Zheng
Earth Syst. Sci. Data, 15, 2235–2258, https://doi.org/10.5194/essd-15-2235-2023, https://doi.org/10.5194/essd-15-2235-2023, 2023
Short summary
Short summary
Climate reconstruction from proxy data can help evaluate climate models. We present pollen-based reconstructions of mean July temperature, mean annual temperature, and annual precipitation from 2594 pollen records from the Northern Hemisphere, using three reconstruction methods (WA-PLS, WA-PLS_tailored, and MAT). Since no global or hemispheric synthesis of quantitative precipitation changes are available for the Holocene so far, this dataset will be of great value to the geoscientific community.
Thomas Kleinen, Sergey Gromov, Benedikt Steil, and Victor Brovkin
Clim. Past, 19, 1081–1099, https://doi.org/10.5194/cp-19-1081-2023, https://doi.org/10.5194/cp-19-1081-2023, 2023
Short summary
Short summary
We modelled atmospheric methane continuously from the last glacial maximum to the present using a state-of-the-art Earth system model. Our model results compare well with reconstructions from ice cores and improve our understanding of a very intriguing period of Earth system history, the deglaciation, when atmospheric methane changed quickly and strongly. Deglacial methane changes are driven by emissions from tropical wetlands, with wetlands in high northern latitudes being secondary.
Manuel Chevalier, Anne Dallmeyer, Nils Weitzel, Chenzhi Li, Jean-Philippe Baudouin, Ulrike Herzschuh, Xianyong Cao, and Andreas Hense
Clim. Past, 19, 1043–1060, https://doi.org/10.5194/cp-19-1043-2023, https://doi.org/10.5194/cp-19-1043-2023, 2023
Short summary
Short summary
Data–data and data–model vegetation comparisons are commonly based on comparing single vegetation estimates. While this approach generates good results on average, reducing pollen assemblages to single single plant functional type (PFT) or biome estimates can oversimplify the vegetation signal. We propose using a multivariate metric, the Earth mover's distance (EMD), to include more details about the vegetation structure when performing such comparisons.
Boris K. Biskaborn, Amy Forster, Gregor Pfalz, Lyudmila A. Pestryakova, Kathleen Stoof-Leichsenring, Jens Strauss, Tim Kröger, and Ulrike Herzschuh
Biogeosciences, 20, 1691–1712, https://doi.org/10.5194/bg-20-1691-2023, https://doi.org/10.5194/bg-20-1691-2023, 2023
Short summary
Short summary
Lake sediment from the Russian Arctic was studied for microalgae and organic matter chemistry dated back to the last glacial 28 000 years. Species and chemistry responded to environmental changes such as the Younger Dryas cold event and the Holocene thermal maximum. Organic carbon accumulation correlated with rates of microalgae deposition only during warm episodes but not during the cold glacial.
Leonore Jungandreas, Cathy Hohenegger, and Martin Claussen
Clim. Past, 19, 637–664, https://doi.org/10.5194/cp-19-637-2023, https://doi.org/10.5194/cp-19-637-2023, 2023
Short summary
Short summary
Increasing the vegetation cover over mid-Holcocene North Africa expands the West African monsoon ∼ 4–5° further north. This northward shift of monsoonal precipitation is caused by interactions of the land surface with large-scale monsoon circulation and the coupling of soil moisture to precipitation. We highlight the importance of considering not only how soil moisture influences precipitation but also how different precipitation characteristics alter the soil hydrology via runoff generation.
Furong Li, Marie-José Gaillard, Xianyong Cao, Ulrike Herzschuh, Shinya Sugita, Jian Ni, Yan Zhao, Chengbang An, Xiaozhong Huang, Yu Li, Hongyan Liu, Aizhi Sun, and Yifeng Yao
Earth Syst. Sci. Data, 15, 95–112, https://doi.org/10.5194/essd-15-95-2023, https://doi.org/10.5194/essd-15-95-2023, 2023
Short summary
Short summary
The objective of this study is present the first gridded and temporally continuous quantitative plant-cover reconstruction for temperate and northern subtropical China over the last 12 millennia. The reconstructions are based on 94 pollen records and include estimates for 27 plant taxa, 10 plant functional types, and 3 land-cover types. The dataset is suitable for palaeoclimate modelling and the evaluation of simulated past vegetation cover and anthropogenic land-cover change from models.
Timon Miesner, Ulrike Herzschuh, Luidmila A. Pestryakova, Mareike Wieczorek, Evgenii S. Zakharov, Alexei I. Kolmogorov, Paraskovya V. Davydova, and Stefan Kruse
Earth Syst. Sci. Data, 14, 5695–5716, https://doi.org/10.5194/essd-14-5695-2022, https://doi.org/10.5194/essd-14-5695-2022, 2022
Short summary
Short summary
We present data which were collected on expeditions to the northeast of the Russian Federation. One table describes the 226 locations we visited during those expeditions, and the other describes 40 289 trees which we recorded at these locations. We found out that important information on the forest cannot be predicted precisely from satellites. Thus, for anyone interested in distant forests, it is important to go to there and take measurements or use data (as presented here).
Femke van Geffen, Birgit Heim, Frederic Brieger, Rongwei Geng, Iuliia A. Shevtsova, Luise Schulte, Simone M. Stuenzi, Nadine Bernhardt, Elena I. Troeva, Luidmila A. Pestryakova, Evgenii S. Zakharov, Bringfried Pflug, Ulrike Herzschuh, and Stefan Kruse
Earth Syst. Sci. Data, 14, 4967–4994, https://doi.org/10.5194/essd-14-4967-2022, https://doi.org/10.5194/essd-14-4967-2022, 2022
Short summary
Short summary
SiDroForest is an attempt to remedy data scarcity regarding vegetation data in the circumpolar region, whilst providing adjusted and labeled data for machine learning and upscaling practices. SiDroForest contains four datasets that include SfM point clouds, individually labeled trees, synthetic tree crowns and labeled Sentinel-2 patches that provide insights into the vegetation composition and forest structure of two important vegetation transition zones in Siberia, Russia.
Mateo Duque-Villegas, Martin Claussen, Victor Brovkin, and Thomas Kleinen
Clim. Past, 18, 1897–1914, https://doi.org/10.5194/cp-18-1897-2022, https://doi.org/10.5194/cp-18-1897-2022, 2022
Short summary
Short summary
Using an Earth system model of intermediate complexity, we quantify contributions of the Earth's orbit, greenhouse gases (GHGs) and ice sheets to the strength of Saharan greening during late Quaternary African humid periods (AHPs). Orbital forcing is found as the dominant factor, having a critical threshold and accounting for most of the changes in the vegetation response. However, results suggest that GHGs may influence the orbital threshold and thus may play a pivotal role for future AHPs.
Ulrike Herzschuh, Chenzhi Li, Thomas Böhmer, Alexander K. Postl, Birgit Heim, Andrei A. Andreev, Xianyong Cao, Mareike Wieczorek, and Jian Ni
Earth Syst. Sci. Data, 14, 3213–3227, https://doi.org/10.5194/essd-14-3213-2022, https://doi.org/10.5194/essd-14-3213-2022, 2022
Short summary
Short summary
Pollen preserved in environmental archives such as lake sediments and bogs are extensively used for reconstructions of past vegetation and climate. Here we present LegacyPollen 1.0, a dataset of 2831 fossil pollen records from all over the globe that were collected from publicly available databases. We harmonized the names of the pollen taxa so that all datasets can be jointly investigated. LegacyPollen 1.0 is available as an open-access dataset.
Ramesh Glückler, Rongwei Geng, Lennart Grimm, Izabella Baisheva, Ulrike Herzschuh, Kathleen R. Stoof-Leichsenring, Stefan Kruse, Andrei Andreev, Luidmila Pestryakova, and Elisabeth Dietze
EGUsphere, https://doi.org/10.5194/egusphere-2022-395, https://doi.org/10.5194/egusphere-2022-395, 2022
Preprint archived
Short summary
Short summary
Despite rapidly intensifying wildfire seasons in Siberian boreal forests, little is known about long-term relationships between changes in vegetation and shifts in wildfire activity. Using lake sediment proxies, we reconstruct such environmental changes over the past 10,800 years in Central Yakutia. We find that a more open forest may facilitate increased amounts of vegetation burning. The present-day dense larch forest might yet be mediating the current climate-driven wildfire intensification.
Nora Farina Specht, Martin Claussen, and Thomas Kleinen
Clim. Past, 18, 1035–1046, https://doi.org/10.5194/cp-18-1035-2022, https://doi.org/10.5194/cp-18-1035-2022, 2022
Short summary
Short summary
Palaeoenvironmental records only provide a fragmentary picture of the lake and wetland extent in North Africa during the mid-Holocene. Therefore, we investigate the possible range of mid-Holocene precipitation changes caused by an estimated small and maximum lake extent and a maximum wetland extent. Results show a particularly strong monsoon precipitation response to lakes and wetlands over the Western Sahara and an increased monsoon precipitation when replacing lakes with vegetated wetlands.
Chenzhi Li, Alexander K. Postl, Thomas Böhmer, Xianyong Cao, Andrew M. Dolman, and Ulrike Herzschuh
Earth Syst. Sci. Data, 14, 1331–1343, https://doi.org/10.5194/essd-14-1331-2022, https://doi.org/10.5194/essd-14-1331-2022, 2022
Short summary
Short summary
Here we present a global chronology framework of 2831 palynological records, including globally harmonized chronologies covering up to 273 000 years. A comparison with the original chronologies reveals a major improvement according to our assessment. Our chronology framework and revised chronologies will interest a broad geoscientific community, as it provides the opportunity to make use in synthesis studies of, for example, pollen-based vegetation and climate change.
Stefan Kruse, Simone M. Stuenzi, Julia Boike, Moritz Langer, Josias Gloy, and Ulrike Herzschuh
Geosci. Model Dev., 15, 2395–2422, https://doi.org/10.5194/gmd-15-2395-2022, https://doi.org/10.5194/gmd-15-2395-2022, 2022
Short summary
Short summary
We coupled established models for boreal forest (LAVESI) and permafrost dynamics (CryoGrid) in Siberia to investigate interactions of the diverse vegetation layer with permafrost soils. Our tests showed improved active layer depth estimations and newly included species growth according to their species-specific limits. We conclude that the new model system can be applied to simulate boreal forest dynamics and transitions under global warming and disturbances, expanding our knowledge.
Jooyeop Lee, Martin Claussen, Jeongwon Kim, Je-Woo Hong, In-Sun Song, and Jinkyu Hong
Clim. Past, 18, 313–326, https://doi.org/10.5194/cp-18-313-2022, https://doi.org/10.5194/cp-18-313-2022, 2022
Short summary
Short summary
It is still a challenge to simulate the so–called Green Sahara (GS), which was a wet and vegetative Sahara region in the mid–Holocene, using current climate models. Our analysis shows that Holocene greening is simulated better if the amount of soil nitrogen and soil texture is properly modified for the humid and vegetative GS period. Future climate simulation needs to consider consequent changes in soil nitrogen and texture with changes in vegetation cover for proper climate simulations.
Anne Dallmeyer, Martin Claussen, Stephan J. Lorenz, Michael Sigl, Matthew Toohey, and Ulrike Herzschuh
Clim. Past, 17, 2481–2513, https://doi.org/10.5194/cp-17-2481-2021, https://doi.org/10.5194/cp-17-2481-2021, 2021
Short summary
Short summary
Using the comprehensive Earth system model, MPI-ESM1.2, we explore the global Holocene vegetation changes and interpret them in terms of the Holocene climate change. The model results reveal that most of the Holocene vegetation transitions seen outside the high northern latitudes can be attributed to modifications in the intensity of the global summer monsoons.
Stuart A. Vyse, Ulrike Herzschuh, Gregor Pfalz, Lyudmila A. Pestryakova, Bernhard Diekmann, Norbert Nowaczyk, and Boris K. Biskaborn
Biogeosciences, 18, 4791–4816, https://doi.org/10.5194/bg-18-4791-2021, https://doi.org/10.5194/bg-18-4791-2021, 2021
Short summary
Short summary
Lakes act as important stores of organic carbon and inorganic sediment material. This study provides a first investigation into carbon and sediment accumulation and storage within an Arctic glacial lake from Far East Russia. It shows that major shifts are related to palaeoclimate variation that affects the development of the lake and its surrounding catchment. Spatial differences to other lake systems from other regions may reflect variability in processes controlled by latitude and altitude.
Leonore Jungandreas, Cathy Hohenegger, and Martin Claussen
Clim. Past, 17, 1665–1684, https://doi.org/10.5194/cp-17-1665-2021, https://doi.org/10.5194/cp-17-1665-2021, 2021
Short summary
Short summary
We investigate the impact of explicitly resolving convection on the mid-Holocene West African Monsoon rain belt by employing the ICON climate model in high resolution. While the spatial distribution and intensity of the precipitation are improved by this technique, the monsoon extents further north and the mean summer rainfall is higher in the simulation with parameterized convection.
Ramesh Glückler, Ulrike Herzschuh, Stefan Kruse, Andrei Andreev, Stuart Andrew Vyse, Bettina Winkler, Boris K. Biskaborn, Luidmila Pestryakova, and Elisabeth Dietze
Biogeosciences, 18, 4185–4209, https://doi.org/10.5194/bg-18-4185-2021, https://doi.org/10.5194/bg-18-4185-2021, 2021
Short summary
Short summary
Data about past fire activity are very sparse in Siberia. This study presents a first high-resolution record of charcoal particles from lake sediments in boreal eastern Siberia. It indicates that current levels of charcoal accumulation are not unprecedented. While a recent increase in reconstructed fire frequency coincides with rising temperatures and increasing human activity, vegetation composition does not seem to be a major driver behind changes in the fire regime in the past two millennia.
Lydia Stolpmann, Caroline Coch, Anne Morgenstern, Julia Boike, Michael Fritz, Ulrike Herzschuh, Kathleen Stoof-Leichsenring, Yury Dvornikov, Birgit Heim, Josefine Lenz, Amy Larsen, Katey Walter Anthony, Benjamin Jones, Karen Frey, and Guido Grosse
Biogeosciences, 18, 3917–3936, https://doi.org/10.5194/bg-18-3917-2021, https://doi.org/10.5194/bg-18-3917-2021, 2021
Short summary
Short summary
Our new database summarizes DOC concentrations of 2167 water samples from 1833 lakes in permafrost regions across the Arctic to provide insights into linkages between DOC and environment. We found increasing lake DOC concentration with decreasing permafrost extent and higher DOC concentrations in boreal permafrost sites compared to tundra sites. Our study shows that DOC concentration depends on the environmental properties of a lake, especially permafrost extent, ecoregion, and vegetation.
Cited articles
Adam, M., Weitzel, N., and Rehfeld, K.: Identifying Global-Scale Patterns of Vegetation Change During the Last Deglaciation From Paleoclimate Networks, Palaeogeogr. Palaeocl., 36, e2021PA004265, https://doi.org/10.1029/2021PA004265, 2021.
Allen, J. R. M., Forrest, M., Hickler, T., Singarayer, J. S., Valdes, P. J., and Huntley, B.: Global vegetation patterns of the past 140,000 years, J. Biogeogr., 47, 2073–2090, https://doi.org/10.1111/jbi.13930, 2020.
Anhuf, D., Ledru, M.-P., Behling, H., Da Cruz, F. W., Cordeiro, R. C., Van der Hammen, T., Karmann, I., Marengo, J. A., De Oliveira, P. E., Pessenda, L., Siffedine, A., Albuquerque, A. L., and Da Silva Dias, P. L.: Paleo-environmental change in Amazonian and African rainforest during the LGM, Palaeogeogr. Palaeocl., 239, 510–527, https://doi.org/10.1016/j.palaeo.2006.01.017, 2006.
Annan, J. D., Hargreaves, J. C., and Mauritsen, T.: A new global surface temperature reconstruction for the Last Glacial Maximum, Clim. Past, 18, 1883–1896, https://doi.org/10.5194/cp-18-1883-2022, 2022.
Arora, V. K., Katavouta, A., Williams, R. G., Jones, C. D., Brovkin, V., Friedlingstein, P., Schwinger, J., Bopp, L., Boucher, O., Cadule, P., Chamberlain, M. A., Christian, J. R., Delire, C., Fisher, R. A., Hajima, T., Ilyina, T., Joetzjer, E., Kawamiya, M., Koven, C. D., Krasting, J. P., Law, R. M., Lawrence, D. M., Lenton, A., Lindsay, K., Pongratz, J., Raddatz, T., Séférian, R., Tachiiri, K., Tjiputra, J. F., Wiltshire, A., Wu, T., and Ziehn, T.: Carbon–concentration and carbon–climate feedbacks in CMIP6 models and their comparison to CMIP5 models, Biogeosciences, 17, 4173–4222, https://doi.org/10.5194/bg-17-4173-2020, 2020.
Bartlein, P. J., Harrison, S. P., Brewer, S., Connor, S., Davis, B. A. S., Gajewski, K., Guiot, J., Harrison-Prentice, T. I., Henderson, A., Peyron, O., Prentice, I. C., Scholze, M., Seppa, H., Shuman, B., Sugita, S., Thompson, R. S., Viau, A. E., Williams, J., and Wu, H.: Pollen-based continental climate reconstructions at 6 and 21 ka: a global synthesis, Clim. Dynam., 37, 775–802, https://doi.org/10.1007/s00382-010-0904-1, 2011.
Batchelor, C. L., Margold, M., Krapp, M., Murton, D. K., Dalton, A. S., Gibbard, P. L., Stokes, C. R., Murton, J. B., and Manica, A.: The configuration of Northern Hemisphere ice sheets through the Quaternary, Nat. Commun., 10, 3713, https://doi.org/10.1038/s41467-019-11601-2, 2019.
Berger, A. L.: Long-term variations of daily insolation and Quaternary climatic changes, Journal of Atmospheric Sciences, 35, 2361–2367, https://doi.org/10.1175/1520-0469(1978)035<2362:LTVODI>2.0.CO;2, 1978.
Beyer, R., Krapp, M., and Manica, A.: An empirical evaluation of bias correction methods for palaeoclimate simulations, Clim. Past, 16, 1493–1508, https://doi.org/10.5194/cp-16-1493-2020, 2020.
Bigelow, N. H., Brubaker, L. B., Edwards, M. E., Harrison, S. P., Prentice, I. C., Anderson, P. M., Andreev, A. A., Bartlein, P. J., Christensen, T. R., Cramer, W., Kaplan, J. O., Lozhkin, A. V., Matveyeva, N. V., Murray, D. F., McGuire, A. D., Razzhivin, V. Y., Ritchie, J. C., Smith, B., Walker, D. A., Gajewski, K., Wolf, V., Holmqvist, B. H., Igarashi, Y., Kremenetskii, K., Paus, A., Pisaric, M. F. J., and Volkova, V. S.: Climate change and Arctic ecosystems: 1. Vegetation changes north of 55° N between the last glacial maximum, mid-Holocene, and present, J. Geophys. Res.-Atmos., 108, https://doi.org/10.1029/2002jd002558, 2003.
Binney, H., Edwards, M., Macias-Fauria, M., Lozhkin, A., Anderson, P., Kaplan, J. O., Andreev, A., Bezrukova, E., Blyakharchuk, T., Jankovska, V., Khazina, I., Krivonogov, S., Kremenetski, K., Nield, J., Novenko, E., Ryabogina, N., Solovieva, N., Willis, K., and Zernitskaya, V.: Vegetation of Eurasia from the last glacial maximum to present: Key biogeographic patterns, Quaternary Sci. Rev., 157, 80–97, https://doi.org/10.1016/j.quascirev.2016.11.022, 2017.
Braconnot, P., Harrison, S. P., Kageyama, M., Bartlein, P. J., Masson-Delmotte, V., Abe-Ouchi, A., Otto-Bliesner, B., and Zhao, Y.: Evaluation of climate models using palaeoclimatic data, Nat. Clim. Change, 2, 417–424, https://doi.org/10.1038/NCLIMATE1456, 2012.
Cao, X., Tian, F., Li, F., Gaillard, M.-J., Rudaya, N., Xu, Q., and Herzschuh, U.: Pollen-based quantitative land-cover reconstruction for northern Asia covering the last 40 ka cal BP, Clim. Past, 15, 1503–1536, https://doi.org/10.5194/cp-15-1503-2019, 2019a.
Cao, X., Tian, F., Li, F., Gaillard, M.-J., Rudaya, N., Xu, Q., and Herzschuh, U.: Pollen-based quantitative land-cover reconstruction for northern Asia covering the last 40 ka cal BP, Clim. Past, 15, 1503–1536, https://doi.org/10.5194/cp-15-1503-2019, 2019b.
Charrier, G., Nolf, M., Leitinger, G., Charra-Vaskou, K., Losso, A., Tappeiner, U., Améglio, T., and Mayr, S.: Monitoring of Freezing Dynamics in Trees: A Simple Phase Shift Causes Complexity, Plant Physiol., 173, 2196–2207, https://doi.org/10.1104/pp.16.01815, 2017.
Cheng, J., Wu, H., Liu, Z., Gu, P., Wang, J., Zhao, C., Li, Q., Chen, H., Lu, H., Hu, H., Gao, Y., Yu, M., and Song, Y.: Vegetation feedback causes delayed ecosystem response to East Asian Summer Monsoon Rainfall during the Holocene, Nat. Commun., 12, 1–9, https://doi.org/10.1038/s41467-021-22087-2, 2021.
Clark, P. U., Shakun, J. D., Baker, P. A., Bartlein, P. J., Brewer, S., Brook, E., Carlson, A. E., Cheng, H., Kaufman, D. S., Liu, Z., Marchitto, T. M., Mix, A. C., Morrill, C., Otto-Bliesner, B. L., Pahnke, K., Russell, J. M., Whitlock, C., Adkins, J. F., Blois, J. L., Clark, J., Colman, S. M., Curry, W. B., Flower, B. P., He, F., Johnson, T. C., Lynch-Stieglitz, J., Markgraf, V., McManus, J., Mitrovica, J. X., Moreno, P. I., and Williams, J. W.: Global climate evolution during the last deglaciation, P. Natl. Acad. Sci. USA, 109, E1134–E1142, https://doi.org/10.1073/pnas.1116619109, 2012.
Claussen, M., Selent, K., Brovkin, V., Raddatz, T., and Gayler, V.: Impact of CO2 and climate on Last Glacial maximum vegetation – a factor separation, Biogeosciences, 10, 3593–3604, https://doi.org/10.5194/bg-10-3593-2013, 2013.
Dallmeyer, A.: From Ice Age to Present: Tree Cover Dynamics in REVEALS and MPI-ESM (Video), Edmond, V1 [video], https://doi.org/10.17617/3.O2GGHG, 2026.
Dallmeyer, A. and Weitzel, N.: Processed Data and Scripts for “Unravelling the Tree Cover Dynamics over the Last 20,000 Years on the Northern Hemisphere”, Edmond [data set], https://doi.org/10.17617/3.HHMFGX, 2026.
Dallmeyer, A., Claussen, M., and Brovkin, V.: Harmonising plant functional type distributions for evaluating Earth system models, Clim. Past, 15, 335–366, https://doi.org/10.5194/cp-15-335-2019, 2019.
Dallmeyer, A., Claussen, M., Lorenz, S. J., Sigl, M., Toohey, M., and Herzschuh, U.: Holocene vegetation transitions and their climatic drivers in MPI-ESM1.2, Clim. Past, 17, 2481–2513, https://doi.org/10.5194/cp-17-2481-2021, 2021.
Dallmeyer, A., Kleinen, T., Claussen, M., Weitzel, N., Cao, X., and Herzschuh, U.: The deglacial forest conundrum, Nat. Commun., 13, 6035, https://doi.org/10.1038/s41467-022-33646-6, 2022.
Dallmeyer, A., Poska, A., Marquer, L., Seim, A., and Gaillard, M.-J.: The challenge of comparing pollen-based quantitative vegetation reconstructions with outputs from vegetation models – a European perspective, Clim. Past, 19, 1531–1557, https://doi.org/10.5194/cp-19-1531-2023, 2023.
Davis, B. A. S., Fasel, M., Kaplan, J. O., Russo, E., and Burke, A.: The climate and vegetation of Europe, northern Africa, and the Middle East during the Last Glacial Maximum (21 000 yr BP) based on pollen data, Clim. Past, 20, 1939–1988, https://doi.org/10.5194/cp-20-1939-2024, 2024.
Dawson, A., Williams, J. W., Gaillard, M.-J., Goring, S. J., Pirzamanbein, B., Lindstrom, J., Anderson, R. S., Brunelle, A., Foster, D., Gajewski, K., Gavin, D. G., Lacourse, T., Minckley, T. A., Oswald, W., Shuman, B., and Whitlock, C.: Holocene land cover change in North America: continental trends, regional drivers, and implications for vegetation–atmosphere feedbacks, Clim. Past, 21, 2031–2060, https://doi.org/10.5194/cp-21-2031-2025, 2025.
DiMiceli, D., Sohlberg, R., and Townshend, J.: MODIS/Terra Vegetation Continuous Fields Yearly L3 Global 250m SIN Grid V061, NASA Land Processes Distributed Active Archive Center [data set], https://doi.org/10.5067/MODIS/MOD44B.061, 2022.
Dupont, L. M., Jahns, S., Marret, F., and Ning, S.: Vegetation change in equatorial West Africa: time-slices for the last 150 ka, Palaeogeogr. Palaeocl., 155, 95–122, https://doi.org/10.1016/S0031-0182(99)00095-4, 2000.
Ellis, E. C., Kaplan, J. O., Fuller, D. Q., Vavrus, S., Klein Goldewijk, K., and Verburg, P. H.: Used planet: A global history, P. Natl. Acad. Sci. USA, 110, 7978–7985, https://doi.org/10.1073/pnas.1217241110, 2013.
Fyfe, R. M., de Beaulieu, J.-L., Binney, H., Bradshaw, R. H. W., Brewer, S., Le Flao, A., Finsinger, W., Gaillard, M.-J., Giesecke, T., Gil-Romera, G., Grimm, E. C., Huntley, B., Kunes, P., Kühl, N., Leydet, M., Lotter, A. F., Tarasov, P. E., and Tonkov, S.: The European Pollen Database: past efforts and current activities, Veg. Hist. Archaeobot., 18, 417–424, https://doi.org/10.1007/s00334-009-0215-9, 2009.
Gaillard, M.-J., Sugita, S., Mazier, F., Trondman, A.-K., Broström, A., Hickler, T., Kaplan, J. O., Kjellström, E., Kokfelt, U., Kuneš, P., Lemmen, C., Miller, P., Olofsson, J., Poska, A., Rundgren, M., Smith, B., Strandberg, G., Fyfe, R., Nielsen, A. B., Alenius, T., Balakauskas, L., Barnekow, L., Birks, H. J. B., Bjune, A., Björkman, L., Giesecke, T., Hjelle, K., Kalnina, L., Kangur, M., van der Knaap, W. O., Koff, T., Lagerås, P., Latałowa, M., Leydet, M., Lechterbeck, J., Lindbladh, M., Odgaard, B., Peglar, S., Segerström, U., von Stedingk, H., and Seppä, H.: Holocene land-cover reconstructions for studies on land cover-climate feedbacks, Clim. Past, 6, 483–499, https://doi.org/10.5194/cp-6-483-2010, 2010.
Giesecke, T., Ammann, B., and Brande, A.: Palynological richness and evenness: insights from the taxa accumulation curve, Veg. Hist. Archaeobot., 23, 217–228, https://doi.org/10.1007/s00334-014-0435-5, 2014.
Githumbi, E., Fyfe, R., Gaillard, M.-J., Trondman, A.-K., Mazier, F., Nielsen, A.-B., Poska, A., Sugita, S., Woodbridge, J., Azuara, J., Feurdean, A., Grindean, R., Lebreton, V., Marquer, L., Nebout-Combourieu, N., Stančikaitė, M., Tanţău, I., Tonkov, S., Shumilovskikh, L., and LandClimII data contributors: European pollen-based REVEALS land-cover reconstructions for the Holocene: methodology, mapping and potentials, Earth Syst. Sci. Data, 14, 1581–1619, https://doi.org/10.5194/essd-14-1581-2022, 2022.
Gosling, W. D., Miller, C. S., Shanahan, T. M., Holden, P. B., Overpeck, J. T., and van Langevelde, F.: A stronger role for long-term moisture change than for CO2 in determining tropical woody vegetation change, Science, 376, 653–656, https://doi.org/10.1126/science.abg4618, 2022.
Hao, S., Zhang, X., Duan, Y., Gowan, E. J., Zhu, J., Cauquoin, A., Chen, J., Werner, M., and Chen, F.: Model seasonal and proxy spatial biases revealed by assimilated mid-Holocene seasonal temperatures, Sci. Bull., 70, 2014–2022, https://doi.org/10.1016/j.scib.2025.03.039, 2025.
Harris, I., Osborn, T. J., Jones, P., and Lister, D.: Version 4 of the CRU TS monthly high-resolution gridded multivariate climate dataset, Scientific Data, 7, 109, https://doi.org/10.1038/s41597-020-0453-3, 2020.
Harrison, S. P., Bartlein, P. J., Izumi, K., Li, G., Annan, J., Hargreaves, J., Braconnot, P., and Kageyama, M.: Evaluation of CMIP5 palaeo-simulations to improve climate projections, Nat. Clim. Change, 5, 735–743, https://doi.org/10.1038/nclimate2649, 2015.
Helbig, M., Pappas, C., and Sonnentag, O.: Permafrost thaw and wildfire: Equally important drivers of boreal tree cover changes in the Taiga Plains, Canada, Geophys. Res. Lett., 43, 1598–1606, https://doi.org/10.1002/2015GL067193, 2016.
Herzschuh, U., Birks, H. J. B., Laepple, T., Andreev, A., Melles, M., and Brigham-Grette, J.: Glacial legacies on interglacial vegetation at the Pliocene-Pleistocene transition in NE Asia, Nat. Commun., 7, 1–11, https://doi.org/10.1038/ncomms11967, 2016.
Herzschuh, U., Li, C., Böhmer, T., Postl, A. K., Heim, B., Andreev, A. A., Cao, X., Wieczorek, M., and Ni, J.: LegacyPollen 1.0: a taxonomically harmonized global late Quaternary pollen dataset of 2831 records with standardized chronologies, Earth Syst. Sci. Data, 14, 3213–3227, https://doi.org/10.5194/essd-14-3213-2022, 2022.
Horvath, P., Tang, H., Halvorsen, R., Stordal, F., Tallaksen, L. M., Berntsen, T. K., and Bryn, A.: Improving the representation of high-latitude vegetation distribution in dynamic global vegetation models, Biogeosciences, 18, 95–112, https://doi.org/10.5194/bg-18-95-2021, 2021.
Ivanovic, R. F., Gregoire, L. J., Kageyama, M., Roche, D. M., Valdes, P. J., Burke, A., Drummond, R., Peltier, W. R., and Tarasov, L.: Transient climate simulations of the deglaciation 21–9 thousand years before present (version 1) – PMIP4 Core experiment design and boundary conditions, Geosci. Model Dev., 9, 2563–2587, https://doi.org/10.5194/gmd-9-2563-2016, 2016.
Jin, H., Huang, Y., Bense, V. F., Ma, Q., Marchenko, S. S., Shepelev, V. V., Hu, Y., Liang, S., Spektor, V. V., Jin, X., Li, X., and Li, X.: Permafrost Degradation and Its Hydrogeological Impacts, Water, 14, 372, https://doi.org/10.3390/w14030372, 2022.
Jin, X.-Y., Jin, H.-J., Iwahana, G., Marchenko, S. S., Luo, D.-L., Li, X.-Y., and Liang, S.-H.: Impacts of climate-induced permafrost degradation on vegetation: A review, Advances in Climate Change Research, 12, 29–47, https://doi.org/10.1016/j.accre.2020.07.002, 2021.
Kaplan, J. O., Krumhardt, K. M., Ellis, E. C., Ruddiman, W. F., Lemmen, C., and Goldewijk, K. K.: Holocene carbon emissions as a result of anthropogenic land cover change, The Holocene, 21, 775–791, https://doi.org/10.1177/0959683610386983, 2011.
Kaplan, J. O., Krumhardt, K. M., Gaillard, M.-J., Sugita, S., Trondman, A.-K., Fyfe, R., Marquer, L., Mazier, F., and Nielsen, A. B.: Constraining the Deforestation History of Europe: Evaluation of Historical Land Use Scenarios with Pollen-Based Land Cover Reconstructions, Land, 6, 91, https://doi.org/10.3390/land6040091, 2017.
Kern, O. A., Maier, A., and Vercauteren, N.: Landscape reconstructions for Europe during the late Last Glacial (60–20 ka BP): a pollen-based REVEALS approach, Earth Syst. Sci. Data, 17, 5997–6023, https://doi.org/10.5194/essd-17-5997-2025, 2025.
Kim, I.-W., Timmermann, A., Kim, J.-E., Rodgers, K. B., Lee, S.-S., Lee, H., and Wieder, W. R.: Abrupt increase in Arctic-Subarctic wildfires caused by future permafrost thaw, Nat. Commun., 15, 7868, https://doi.org/10.1038/s41467-024-51471-x, 2024.
Kleinen, T., Gromov, S., Steil, B., and Brovkin, V.: Atmospheric methane since the last glacial maximum was driven by wetland sources, Clim. Past, 19, 1081–1099, https://doi.org/10.5194/cp-19-1081-2023, 2023a.
Kleinen, T., Gromov, S., Steil, B., and Brovkin, V.: PalMod2 MPI-M MPI-ESM1-2-CR-CH4 transient-deglaciation-prescribed-glac1d-methane, World Data Center for Climate (WDCC) at DKRZ [data set], https://doi.org/10.26050/WDCC/PMMXMCHTD, 2023b.
Köhler, P., Nehrbass-Ahles, C., Schmitt, J., Stocker, T. F., and Fischer, H.: A 156 kyr smoothed history of the atmospheric greenhouse gases CO2, CH4, and N2O and their radiative forcing, Earth Syst. Sci. Data, 9, 363–387, https://doi.org/10.5194/essd-9-363-2017, 2017.
Kruse, S., Stuenzi, S. M., Boike, J., Langer, M., Gloy, J., and Herzschuh, U.: Novel coupled permafrost–forest model (LAVESI–CryoGrid v1.0) revealing the interplay between permafrost, vegetation, and climate across eastern Siberia, Geosci. Model Dev., 15, 2395–2422, https://doi.org/10.5194/gmd-15-2395-2022, 2022.
Leroy, S. A. G., Arpe, K., Mikolajewicz, U., and Wu, J.: Climate simulations and pollen data reveal the distribution and connectivity of temperate tree populations in eastern Asia during the Last Glacial Maximum, Clim. Past, 16, 2039–2054, https://doi.org/10.5194/cp-16-2039-2020, 2020.
Li, C., Dallmeyer, A., Ni, J., Chevalier, M., Willeit, M., Andreev, A. A., Cao, X., Schild, L., Heim, B., Wieczorek, M., and Herzschuh, U.: Global biome changes over the last 21 000 years inferred from model–data comparisons, Clim. Past, 21, 1001–1024, https://doi.org/10.5194/cp-21-1001-2025, 2025a.
Li, C., Ni, J., Böhmer, T., Cao, X., Zhou, B., Liao, M., Li, K., Schild, L., Wieczorek, M., Heim, B., and Herzschuh, U.: LegacyPollen2.0: an updated global taxonomically and temporally standardized fossil pollen dataset of 3680 palynological records, PANGAEA [data set], https://doi.org/10.1594/PANGAEA.965907, 2025b.
Li, F., Gaillard, M.-J., Cao, X., Herzschuh, U., Sugita, S., Ni, J., Zhao, Y., An, C., Huang, X., Li, Y., Liu, H., Sun, A., and Yao, Y.: Gridded pollen-based Holocene regional plant cover in temperate and northern subtropical China suitable for climate modelling, Earth Syst. Sci. Data, 15, 95–112, https://doi.org/10.5194/essd-15-95-2023, 2023a.
Li, F., Gaillard, M.-J., Cao, X., Herzschuh, U., Sugita, S., Ni, J., Zhao, Y., An, C., Huang, X., Li, Y., Liu, H., Sun, A., and Yao, Y.: Gridded pollen-based Holocene regional plant cover in temperate and northern subtropical China suitable for climate modelling, Earth Syst. Sci. Data, 15, 95–112, https://doi.org/10.5194/essd-15-95-2023, 2023b.
Li, G., Chen, W., Zhang, X., Yang, Z., Wang, Z., and Bi, P.: Spatiotemporal changes and driving factors of vegetation in 14 different climatic regions in the global from 1981 to 2018, Environ. Sci. Pollut. Res., 29, 75322–75337, https://doi.org/10.1007/s11356-022-21138-5, 2022a.
Li, W., Tian, F., Rudaya, N., Herzschuh, U., and Cao, X.: Pollen-Based Holocene Thawing-History of Permafrost in Northern Asia and Its Potential Impacts on Climate Change, Front. Ecol. Evol., 10, https://doi.org/10.3389/fevo.2022.894471, 2022b.
MacDonald, G. M., Velichko, A. A., Kremenetski, C. V., Borisova, O. K., Goleva, A. A., Andreev, A. A., Cwynar, L. C., Riding, R. T., Forman, S. L., Edwards, T. W. D., Aravena, R., Hammarlund, D., Szeicz, J. M., and Gattaulin, V. N.: Holocene Treeline History and Climate Change Across Northern Eurasia, Quaternary Res., 53, 302–311, https://doi.org/10.1006/qres.1999.2123, 2000.
Maraun, D. and Widmann, M.: Statistical Downscaling and Bias Correction for Climate Research, 1st Edn., Cambridge University Press, https://doi.org/10.1017/9781107588783, 2018.
Marquer, L., Gaillard, M.-J., Sugita, S., Poska, A., Trondman, A.-K., Mazier, F., Nielsen, A. B., Fyfe, R. M., Jönsson, A. M., Smith, B., Kaplan, J. O., Alenius, T., Birks, H. J. B., Bjune, A. E., Christiansen, J., Dodson, J., Edwards, K. J., Giesecke, T., Herzschuh, U., Kangur, M., Koff, T., Latałowa, M., Lechterbeck, J., Olofsson, J., and Seppä, H.: Quantifying the effects of land use and climate on Holocene vegetation in Europe, Quaternary Sci. Rev., 171, 20–37, https://doi.org/10.1016/j.quascirev.2017.07.001, 2017.
Mauritsen, T., Bader, J., Becker, T., Behrens, J., Bittner, M., Brokopf, R., Brovkin, V., Claussen, M., Crueger, T., Esch, M., Fast, I., Fiedler, S., Fläschner, D., Gayler, V., Giorgetta, M., Goll, D. S., Haak, H., Hagemann, S., Hedemann, C., Hohenegger, C., Ilyina, T., Jahns, T., Jimenéz-de-la-Cuesta, D., Jungclaus, J., Kleinen, T., Kloster, S., Kracher, D., Kinne, S., Kleberg, D., Lasslop, G., Kornblueh, L., Marotzke, J., Matei, D., Meraner, K., Mikolajewicz, U., Modali, K., Möbis, B., Müller, W. A., Nabel, J. E. M. S., Nam, C. C. W., Notz, D., Nyawira, S., Paulsen, H., Peters, K., Pincus, R., Pohlmann, H., Pongratz, J., Popp, M., Raddatz, T. J., Rast, S., Redler, R., Reick, C. H., Rohrschneider, T., Schemann, V., Schmidt, H., Schnur, R., Schulzweida, U., Six, K. D., Stein, L., Stemmler, I., Stevens, B., Storch, J., Tian, F., Voigt, A., Vrese, P., Wieners, K., Wilkenskjeld, S., Winkler, A., and Roeckner, E.: Developments in the MPI-M Earth System Model version 1.2 (MPI-ESM1.2) and Its Response to Increasing CO 2, J. Adv. Model. Earth Syst., 11, 998–1038, https://doi.org/10.1029/2018MS001400, 2019.
McGee, D.: Glacial–Interglacial Precipitation Changes, Annu. Rev. Mar. Sci., 12, 525–557, https://doi.org/10.1146/annurev-marine-010419-010859, 2020.
Montesano, P. M., Nelson, R., Sun, G., Margolis, H., Kerber, A., and Ranson, K. J.: MODIS tree cover validation for the circumpolar taiga–tundra transition zone, Remote Sens. Environ., 113, 2130–2141, https://doi.org/10.1016/j.rse.2009.05.021, 2009.
Mottl, O., Flantua, S. G. A., Bhatta, K. P., Felde, V. A., Giesecke, T., Goring, S., Grimm, E. C., Haberle, S., Hooghiemstra, H., Ivory, S., Kuneš, P., Wolters, S., Seddon, A. W. R., and Williams, J. W.: Global acceleration in rates of vegetation change over the past 18,000 years, Science, 372, 860–864, https://doi.org/10.1126/science.abg1685, 2021.
Nolan, C., Overpeck, J. T., Allen, J. R. M., Anderson, P. M., Betancourt, J. L., Binney, H. A., Brewer, S., Bush, M. B., Chase, B. M., Cheddadi, R., Djamali, M., Dodson, J., Edwards, M. E., Gosling, W. D., Haberle, S., Hotchkiss, S. C., Huntley, B., Ivory, S. J., Kershaw, A. P., Kim, S. H., Latorre, C., Leydet, M., Lézine, A. M., Liu, K. B., Liu, Y., Lozhkin, A. V., McGlone, M. S., Marchant, R. A., Momohara, A., Moreno, P. I., Müller, S., Otto-Bliesner, B. L., Shen, C., Stevenson, J., Takahara, H., Tarasov, P. E., Tipton, J., Vincens, A., Weng, C., Xu, Q., Zheng, Z., and Jackson, S. T.: Past and future global transformation of terrestrial ecosystems under climate change, Science, 361, 920–923, https://doi.org/10.1126/science.aan5360, 2018.
Osman, M. B., Tierney, J. E., Zhu, J., Tardif, R., Hakim, G. J., King, J., and Poulsen, C. J.: Globally resolved surface temperatures since the Last Glacial Maximum, Nature, 599, 239–244, https://doi.org/10.1038/s41586-021-03984-4, 2021.
Prentice, I. C., Jolly, D., and BIOME 6000 participants: Mid-Holocene and glacial-maximum vegetation geography of the northern continents and Africa, J. Biogeogr., 27, 507–519, https://doi.org/10.1046/j.1365-2699.2000.00425.x, 2000.
Reick, C. H., Raddatz, T., Brovkin, V., and Gayler, V.: Representation of natural and anthropogenic land cover change in MPI-ESM, J. Adv. Model. Earth Syst., 5, 459–482, https://doi.org/10.1002/jame.20022, 2013.
Reick, C. H., Gayler, V., Goll, D., Hagemann, S., Heidkamp, M., Nabel, J. E. M. S., Raddatz, T., Roeckner, E., Schnur, R., and Wilkenskjeld, S.: JSBACH 3 – The land component of the MPI Earth System Model: documentation of version 3.2, Berichte zur Erdsystemforschung, 240, https://doi.org/10.17617/2.3279802, 2021.
Roberts, N., Fyfe, R. M., Woodbridge, J., Gaillard, M.-J., Davis, B. A. S., Kaplan, J. O., Marquer, L., Mazier, F., Nielsen, A. B., Sugita, S., Trondman, A.-K., and Leydet, M.: Europe's lost forests: a pollen-based synthesis for the last 11,000 years, Sci. Rep., 8, 716, https://doi.org/10.1038/s41598-017-18646-7, 2018.
Roebroek, C. T. J., Caporaso, L., Duveiller, G., Davin, E. L., Seneviratne, S. I., and Cescatti, A.: Potential tree cover under current and future climate scenarios, Scientific Data, 12, 564, https://doi.org/10.1038/s41597-025-04408-y, 2025.
Rogers, A., Medlyn, B. E., Dukes, J. S., Bonan, G., von Caemmerer, S., Dietze, M. C., Kattge, J., Leakey, A. D. B., Mercado, L. M., Niinemets, Ü., Prentice, I. C., Serbin, S. P., Sitch, S., Way, D. A., and Zaehle, S.: A roadmap for improving the representation of photosynthesis in Earth system models, New Phytol., 213, 22–42, https://doi.org/10.1111/nph.14283, 2017.
Sakai, A. and Larcher, W.: Frost Survival of Plants, Springer, Berlin, Heidelberg, https://doi.org/10.1007/978-3-642-71745-1, 1987.
Sato, H., Kelley, D. I., Mayor, S. J., Martin Calvo, M., Cowling, S. A., and Prentice, I. C.: Dry corridors opened by fire and low CO2 in Amazonian rainforest during the Last Glacial Maximum, Nat. Geosci., 14, 578–585, https://doi.org/10.1038/s41561-021-00777-2, 2021.
Schild, L.: Northern Hemisphere REVEALS-based forest cover for the past 20 k years, Zenodo [data set], https://doi.org/10.5281/zenodo.17950309, 2025.
Schild, L., Ewald, P., Li, C., Hébert, R., Laepple, T., and Herzschuh, U.: LegacyVegetation: Northern Hemisphere reconstruction of past plant cover and total tree cover from pollen archives of the last 14 kyr, Earth Syst. Sci. Data, 17, 2007–2033, https://doi.org/10.5194/essd-17-2007-2025, 2025.
Seddon, A. W. R., Macias-Fauria, M., Long, P. R., Benz, D., and Willis, K. J.: Sensitivity of global terrestrial ecosystems to climate variability, Nature, 531, 229–232, https://doi.org/10.1038/nature16986, 2016.
Shakun, J. D., Clark, P. U., He, F., Marcott, S. A., Mix, A. C., Liu, Z., Otto-Bliesner, B., Schmittner, A., and Bard, E.: Global warming preceded by increasing carbon dioxide concentrations during the last deglaciation, Nature, 484, 49–54, https://doi.org/10.1038/nature10915, 2012.
Shi, X., Werner, M., Yang, H., D'Agostino, R., Liu, J., Yang, C., and Lohmann, G.: Unraveling the complexities of the Last Glacial Maximum climate: the role of individual boundary conditions and forcings, Clim. Past, 19, 2157–2175, https://doi.org/10.5194/cp-19-2157-2023, 2023.
Stuenzi, S. M., Boike, J., Gädeke, A., Herzschuh, U., Kruse, S., Pestryakova, L. A., Westermann, S., and Langer, M.: Sensitivity of ecosystem-protected permafrost under changing boreal forest structures, Environ. Res. Lett., 16, 084045, https://doi.org/10.1088/1748-9326/ac153d, 2021.
Sugita, S.: Theory of quantitative reconstruction of vegetation I: pollen from large sites REVEALS regional vegetation composition, The Holocene, 17, 229–241, https://doi.org/10.1177/0959683607075837, 2007.
Sweeney, L., Harrison, S. P., and Vander Linden, M.: European tree cover during the Holocene reconstructed from pollen records, Biogeosciences, 22, 4903–4922, https://doi.org/10.5194/bg-22-4903-2025, 2025.
Tarasov, L., Dyke, A. S., Neal, R. M., and Peltier, W. R.: A data-calibrated distribution of deglacial chronologies for the North American ice complex from glaciological modeling, Earth Planet. Sc. Lett., 315–316, 30–40, https://doi.org/10.1016/j.epsl.2011.09.010, 2012.
Theuerkauf, M., Couwenberg, J., Kuparinen, A., and Liebscher, V.: A matter of dispersal: REVEALSinR introduces state-of-the-art dispersal models to quantitative vegetation reconstruction, Veg. Hist. Archaeobot., 25, 541–553, https://doi.org/10.1007/s00334-016-0572-0, 2016.
Thompson, A. J., Zhu, J., Poulsen, C. J., Tierney, J. E., and Skinner, C. B.: Northern Hemisphere vegetation change drives a Holocene thermal maximum, Science Advances, 8, eabj6535, https://doi.org/10.1126/sciadv.abj6535, 2022.
Tian, F., Cao, X., Dallmeyer, A., Lohmann, G., Zhang, X., Ni, J., Andreev, A., Anderson, P. M., Lozhkin, A. V., Bezrukova, E., Rudaya, N., Xu, Q., and Herzschuh, U.: Biome changes and their inferred climatic drivers in northern and eastern continental Asia at selected times since 40 cal ka bp, Veg. Hist. Archaeobot., 27, 365–379, https://doi.org/10.1007/s00334-017-0653-8, 2018.
Tian, Z. and Jiang, D.: Mid-Holocene ocean and vegetation feedbacks over East Asia, Clim. Past, 9, 2153–2171, https://doi.org/10.5194/cp-9-2153-2013, 2013.
Tierney, J. E., Zhu, J., King, J., Malevich, S. B., Hakim, G. J., and Poulsen, C. J.: Glacial cooling and climate sensitivity revisited, Nature, 584, 569, https://doi.org/10.1038/s41586-020-2617-x, 2020.
Trondman, A. -K., Gaillard, M. -J., Mazier, F., Sugita, S., Fyfe, R., Nielsen, A. B., Twiddle, C., Barratt, P., Birks, H. J. B., Bjune, A. E., Björkman, L., Broström, A., Caseldine, C., David, R., Dodson, J., Dörfler, W., Fischer, E., Geel, B., Giesecke, T., Hultberg, T., Kalnina, L., Kangur, M., Knaap, P., Koff, T., Kuneš, P., Lagerås, P., Latałowa, M., Lechterbeck, J., Leroyer, C., Leydet, M., Lindbladh, M., Marquer, L., Mitchell, F. J. G., Odgaard, B. V., Peglar, S. M., Persson, T., Poska, A., Rösch, M., Seppä, H., Veski, S., and Wick, L.: Pollen-based quantitative reconstructions of Holocene regional vegetation cover (plant-functional types and land-cover types) in Europe suitable for climate modelling, Glob. Change Biol., 21, 676–697, https://doi.org/10.1111/gcb.12737, 2015.
Tucker, C. J., Slayback, D. A., Pinzon, J. E., Los, S. O., Myneni, R. B., and Taylor, M. G.: Higher northern latitude normalized difference vegetation index and growing season trends from 1982 to 1999, Int. J. Biometeorol., 45, 184–190, https://doi.org/10.1007/s00484-001-0109-8, 2001.
University of East Anglia Climatic Research Unit, Harris, I. C., Jones, P. D., and Osborn, T.: CRU TS4.08: Climatic Research Unit (CRU) Time-Series (TS) version 4.08 of high-resolution gridded data of month-by-month variation in climate (Jan. 1901–Dec. 2023), NERC EDS Centre for Environmental Data Analysis, https://catalogue.ceda.ac.uk/uuid/715abce1604a42f396f81db83aeb2a4b (last access: 20 January 2026), 2024.
Viereck, L. A.: Wildfire in the Taiga of Alaska, Quaternary Res., 3, 465–495, https://doi.org/10.1016/0033-5894(73)90009-4, 1973.
Wang, X., Edwards, R. L., Auler, A. S., Cheng, H., Kong, X., Wang, Y., Cruz, F. W., Dorale, J. A., and Chiang, H.-W.: Hydroclimate changes across the Amazon lowlands over the past 45,000 years, Nature, 541, 204–207, https://doi.org/10.1038/nature20787, 2017.
Williams, J. W., Shuman, B. N., Webb III, T., Bartlein, P. J., and Leduc, P. L.: Late-Quaternary Vegetation Dynamics in North America: Scaling from Taxa to Biomes, Ecol. Monogr., 74, 309–334, https://doi.org/10.1890/02-4045, 2004.
Williams, J. W., Grimm, E. C., Blois, J. L., Charles, D. F., Davis, E. B., Goring, S. J., Graham, R. W., Smith, A. J., Anderson, M., Arroyo-Cabrales, J., Ashworth, A. C., Betancourt, J. L., Bills, B. W., Booth, R. K., Buckland, P. I., Curry, B. B., Giesecke, T., Jackson, S. T., Latorre, C., Nichols, J., Purdum, T., Roth, R. E., Stryker, M., and Takahara, H.: The Neotoma Paleoecology Database, a multiproxy, international, community-curated data resource, Quaternary Res., 89, 156–177, https://doi.org/10.1017/qua.2017.105, 2018.
Woillez, M.-N., Kageyama, M., Krinner, G., de Noblet-Ducoudré, N., Viovy, N., and Mancip, M.: Impact of CO2 and climate on the Last Glacial Maximum vegetation: results from the ORCHIDEE/IPSL models, Clim. Past, 7, 557–577, https://doi.org/10.5194/cp-7-557-2011, 2011.
Wood, S. N.: Low-Rank Scale-Invariant Tensor Product Smooths for Generalized Additive Mixed Models, Biometrics, 62, 1025–1036, https://doi.org/10.1111/j.1541-0420.2006.00574.x, 2006.
Wood, S. N.: Generalized additive models: an introduction with R, Second edition, CRC Press, Taylor & Francis Group, Boca Raton, FL, 1 pp., https://doi.org/10.1201/9781315370279, 2017.
Wurster, C. M., Bird, M. I., Bull, I. D., Creed, F., Bryant, C., Dungait, J. A. J., and Paz, V.: Forest contraction in north equatorial Southeast Asia during the Last Glacial Period, P. Natl. Acad. Sci. USA, 107, 15508–15511, https://doi.org/10.1073/pnas.1005507107, 2010.
Zhang, P., Luo, Y., Liu, D., Wang, X., and Wang, T.: Pollen-based reconstruction of spatially-explicit vegetation cover over the Tibetan Plateau since the last deglaciation, Earth Syst. Sci. Data, 17, 5557–5570, https://doi.org/10.5194/essd-17-5557-2025, 2025.
Zhang, Y., Renssen, H., Seppä, H., and Valdes, P. J.: Holocene temperature evolution in the Northern Hemisphere high latitudes – Model-data comparisons, Quaternary Sci. Rev., 173, 101–113, https://doi.org/10.1016/j.quascirev.2017.07.018, 2017.
Zhang, Y., Renssen, H., Seppä, H., and Valdes, P. J.: Holocene temperature trends in the extratropical Northern Hemisphere based on inter-model comparisons, J. Quaternary Sci., 33, 464–476, https://doi.org/10.1002/jqs.3027, 2018.
Zhao, A., Li, Z., Zou, L., Wu, J., Stan, K., and Sanchez-Azofeifa, A.: Evaluating dynamic global vegetation models in China: challenges in capturing trends in leaf area and gross primary productivity, Earth Syst. Dynam., 16, 1935–1957, https://doi.org/10.5194/esd-16-1935-2025, 2025.
Zhao, L., Dai, A., and Dong, B.: Changes in global vegetation activity and its driving factors during 1982–2013, Agr. Forest Meteorol., 249, 198–209, https://doi.org/10.1016/j.agrformet.2017.11.013, 2018.
Short summary
We compare pollen-based reconstructions of Northern Hemisphere tree cover over the last 20 000 years with simulations from the Max-Planck-Institute Earth System Model (MPI-ESM). The model captures broad forest trends but misses key regional patterns and the mid-Holocene forest peak. Testing climate drivers reveals mismatches in how temperature, water, and CO2 control forests, pointing to structural limits and the need for improved vegetation processes in models.
We compare pollen-based reconstructions of Northern Hemisphere tree cover over the last 20 000...