NIOZ Royal Netherlands Institute for Sea Research, Department of Marine Geology, P.O. Box 59, 1790 AB Den Burg, Texel, the Netherlands

NIOZ Royal Netherlands Institute for Sea Research, Department of Marine Geology, P.O. Box 59, 1790 AB Den Burg, Texel, the Netherlands

School of Earth and Environment, The University of Western Australia and the UWA Oceans Institute, 35 Stirling Highway, Crawley WA 6009, Australia and the Australian Institute of Marine Science, 39 Fairway, Nedlands WA 6009, Australia

Vrije Universiteit Amsterdam, Faculty of Earth and Life Sciences, De Boelelaan 1105, 1081 HV Amsterdam, the Netherlands

GEOMAR, Helmholtz-Zentrum für Ozeanforschung Kiel, Duesternbrooker Weg 20, 24105 Kiel, Germany

NIOZ Royal Netherlands Institute for Sea Research, Department of Marine Geology, P.O. Box 59, 1790 AB Den Burg, Texel, the Netherlands

University of Amsterdam, Institute for Biodiversity and Ecosystem Dynamics (IBED), Nieuwe Achtergracht 127, 1018 WS Amsterdam, the Netherlands

now at: Unidad Académica de Sistemas Arrecifales, Instituto de Ciencias del Mar y Limnología, Universidad Nacional Autónoma de México, Apdo. Postal 1152, Cancún, Quintana Roo 77500, Mexico

NIOZ Royal Netherlands Institute for Sea Research, Department of Marine Geology, P.O. Box 59, 1790 AB Den Burg, Texel, the Netherlands

Wildlife Conservation Society (WCS), B.P. 8500 Soavimbahoaka, Antananarivo 101, Madagascar

The University of Western Australia and the UWA Oceans Institute, School of Earth and Environment, M004 and ARC Centre of Excellence in Coral Studies, Crawley, Western Australia 6009, Australia

NIOZ Royal Netherlands Institute for Sea Research, Department of Marine Geology, P.O. Box 59, 1790 AB Den Burg, Texel, the Netherlands