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<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:oasis="http://docs.oasis-open.org/ns/oasis-exchange/table" xml:lang="en" dtd-version="3.0" article-type="research-article">
  <front>
    <journal-meta><journal-id journal-id-type="publisher">CP</journal-id><journal-title-group>
    <journal-title>Climate of the Past</journal-title>
    <abbrev-journal-title abbrev-type="publisher">CP</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Clim. Past</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1814-9332</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/cp-22-1203-2026</article-id><title-group><article-title>Shaping the mid-Miocene warmth: a sensitivity study on paleogeography, CO<sub>2</sub> and model physics</article-title><alt-title>Sensitivity of mid-Miocene climate</alt-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1 aff2">
          <name><surname>Renoult</surname><given-names>Martin</given-names></name>
          <email>martin.renoult@geo.su.se</email>
        <ext-link>https://orcid.org/0000-0002-8560-8722</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff2">
          <name><surname>de Boer</surname><given-names>Agatha</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2 aff3">
          <name><surname>Berntell</surname><given-names>Ellen</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-2409-4035</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff2">
          <name><surname>Naik</surname><given-names>Trusha Jagdish</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-9472-0399</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Department of Geological Sciences, Stockholm University, Stockholm, Sweden</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Bolin Center for Climate Research, Stockholm, Sweden</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Department of Meteorology, Stockholm University, Stockholm, Sweden</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Martin Renoult (martin.renoult@geo.su.se)</corresp></author-notes><pub-date><day>17</day><month>June</month><year>2026</year></pub-date>
      
      <volume>22</volume>
      <issue>6</issue>
      <fpage>1203</fpage><lpage>1222</lpage>
      <history>
        <date date-type="received"><day>14</day><month>October</month><year>2025</year></date>
           <date date-type="rev-request"><day>23</day><month>October</month><year>2025</year></date>
           <date date-type="rev-recd"><day>30</day><month>May</month><year>2026</year></date>
           <date date-type="accepted"><day>2</day><month>June</month><year>2026</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2026 Martin Renoult et al.</copyright-statement>
        <copyright-year>2026</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026.html">This article is available from https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026.html</self-uri><self-uri xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026.pdf">The full text article is available as a PDF file from https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d2e130">The mid-Miocene (15.98 to 13.82 Ma) was characterized by substantially warmer temperatures than today and atmospheric CO<sub>2</sub> concentrations comparable to near-future projections. Climate models have generally struggled to reproduce proxy-based reconstructions from this interval, particularly at high latitudes where model temperatures are consistently lower than observations. Here, we present new mid-Miocene simulations using a previously unpublished geography and evaluate the climate's sensitivity to several key components: paleogeography (including land-sea distribution, topography and ice sheets), atmospheric CO<sub>2</sub> concentration, atmospheric model choice, and solar forcing. Our baseline mid-Miocene climate yields a global mean surface temperature (GMST) of 19.8 °C. In mid-Miocene sensitivity experiments of two and four times pre-industrial CO<sub>2</sub> concentrations, consistent with estimates for the mid-Miocene, GMST varies by up to 3.2 °C between simulations. Removal of the Antarctic ice-sheet leads to expected local warming of around 25 °C at the maximum height of the ice sheet, but nevertheless records an overall global cooling of 1.3 °C. Solar forcing and subtle changes of land-sea mask each impact GMST by around 0.2 °C. The choice of atmospheric model substantially affects the simulated mid-Miocene climate through modified feedback mechanisms. We estimate an equilibrium climate sensitivity (ECS) of 2.9 °C (2.5–3.3 °C, 95 % prediction interval) for the mid-Miocene, similar to modern-based estimates from our model (2.8 °C, 2.2–3.4 °C, 95 % prediction interval), indicating the potential for the Miocene to contribute to constraining ECS. Global precipitation is tightly coupled to GMST across all our simulations. As with previous studies, all our simulations, regardless of specific configuration, underestimate high-latitude proxy-reconstructed temperatures. This highlights the need to improve our understanding on polar amplification and on the limitations affecting the proxy record.</p>
  </abstract>
    
<funding-group>
<award-group id="gs1">
<funding-source>Vetenskapsrådet</funding-source>
<award-id>2020-04791</award-id>
</award-group>
</funding-group>
</article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d2e169">The Langhian (15.98 to 13.82 Ma, here referred to as mid-Miocene) is a critical age of the Miocene marked by the warm event known as the mid-Miocene Climatic Optimum (MMCO), when global mean surface temperatures (GMST) were around 7.6 °C warmer than today <xref ref-type="bibr" rid="bib1.bibx69 bib1.bibx22 bib1.bibx10" id="paren.1"/>. During this time, tropical gateways connected the Pacific, Atlantic and Indian oceans, while polar gateways connecting the Arctic with the Atlantic were more restricted than today and the Arctic-Pacific connection was closed off. Flora has substantially changed throughout the Miocene, which also saw the emergence of modern biomes <xref ref-type="bibr" rid="bib1.bibx58" id="paren.2"/>.</p>
      <p id="d2e178">Warm paleoclimates are often investigated as potential analogs for near-future climates <xref ref-type="bibr" rid="bib1.bibx68 bib1.bibx8" id="paren.3"/>. While the analog nature of past climate are never perfect, and there are many differences in how warming manifests itself in past and future climate states  <xref ref-type="bibr" rid="bib1.bibx56 bib1.bibx46" id="paren.4"/>, the past climates offer a comparison of modelling outcomes with proxy data, which in turn can provide insights into climate processes that also act in the future. An example is constraining the important climate change metric of equilibrium climate sensitivity (ECS), the long-term global mean surface temperature change in response to a doubling of CO<sub>2</sub> from its pre-industrial (PI) level, as it was previously done with the Pliocene <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx28 bib1.bibx55 bib1.bibx50 bib1.bibx51" id="paren.5"/>.</p>
      <p id="d2e199">In this context, the mid-Miocene has recently emerged as a particularly exciting testbed for understanding warm climates. In contrast to the more recent interglacials or the older Eocene epoch, mid-Miocene atmospheric CO<sub>2</sub> concentrations are thought to be close to modern values and near-future estimates (400 to 800 ppm), and potentially peaking at much higher values during the MMCO <xref ref-type="bibr" rid="bib1.bibx48 bib1.bibx59" id="paren.6"/>. As a result, this interval has been the focus of several recent modelling and proxy-synthesis studies <xref ref-type="bibr" rid="bib1.bibx58 bib1.bibx10 bib1.bibx44" id="paren.7"/>. A formal two-phase Miocene Model Intercomparison Project (MioMIP) is now under development, with Phase 1 using older boundary conditions and Phase 2 using updated ones. This contracts with MioMIP1, which was as an opportunistic comparison <xref ref-type="bibr" rid="bib1.bibx10" id="paren.8"/>.</p>
      <p id="d2e220">Climate models have underestimated proxy-reconstructed temperatures of the mid-Miocene at equivalent CO<sub>2</sub> concentrations <xref ref-type="bibr" rid="bib1.bibx22 bib1.bibx10" id="paren.9"/>, in particular at high latitudes <xref ref-type="bibr" rid="bib1.bibx30" id="paren.10"><named-content content-type="pre">e.g.</named-content></xref>. This is not a specific issue of the Miocene, as polar amplified warmth is also weak in other warm paleoclimates, such as the Pliocene <xref ref-type="bibr" rid="bib1.bibx28" id="paren.11"/> and the Eocene <xref ref-type="bibr" rid="bib1.bibx39" id="paren.12"/>. This suggests a misrepresentation of high latitude climate feedbacks in models, substantial uncertainties in climate forcings, and/or persistent biases in high-latitude proxies; most likely, it reflects a combination of both model and proxy biases, as suggested for the Pliocene by <xref ref-type="bibr" rid="bib1.bibx64" id="text.13"/>. These points are critical to address, as they can allow to improve simulations of the Miocene, as well as model processes and the accuracy of future model predictions.</p>
      <p id="d2e251">The aim of this study is to explore the mid-Miocene climate and investigate its sensitivity around various uncertain boundary conditions and model parametrizations. We perform simulations with a previously unpublished paleogeography of the mid-Miocene, and compare the simulation to one in the same model using the published paleogeography of <xref ref-type="bibr" rid="bib1.bibx10" id="text.14"/>. We also perform simple computations of Miocene ECS as to evaluate its potential as a new line of evidence for constraining modern-day ECS, in the light of the upcoming MioMIP. We further explore the sensitivity to the Antarctic ice-sheet existence, CO<sub>2</sub> concentrations, solar constant and soil properties. These forcings are actively discussed for MioMIP1 models <xref ref-type="bibr" rid="bib1.bibx10" id="paren.15"/> as well as for the development of MioMIP2, and could be large contributors to the Miocene climate, besides geography. Finally, while not a forcing, we also test the sensitivity of  the mid-Miocene climate to a different atmospheric model, as it is common to have changes of model components between different phases of a model intercomparison project (MIP).</p>

<table-wrap id="T1" specific-use="star"><label>Table 1</label><caption><p id="d2e272">Summary of the experiments carried out in this study. “Getech” refers to the mid-Miocene geography provided by Getech Plc., whereas <xref ref-type="bibr" rid="bib1.bibx10" id="text.16"/> is a mid-Miocene geography. All runs have been integrated for at least 3100 years and up to 5000 years when using CAM5 (Mio_noIS_CAM5), unless specified with <sup>a</sup> where the simulations were ran for 150 years as to estimate ECS. <sup>b</sup>: Solar constant. <sup>c</sup>: Atmospheric model.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="center"/>
     <oasis:colspec colnum="5" colname="col5" align="left"/>
     <oasis:colspec colnum="6" colname="col6" align="left"/>
     <oasis:colspec colnum="7" colname="col7" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Experiment</oasis:entry>
         <oasis:entry colname="col2">Geography</oasis:entry>
         <oasis:entry colname="col3">CO<sub>2</sub> (ppm)</oasis:entry>
         <oasis:entry colname="col4">Sol.<sup>b</sup> (Wm<sup>−2</sup>)</oasis:entry>
         <oasis:entry colname="col5">Ice sheet</oasis:entry>
         <oasis:entry colname="col6">Atm.<sup>c</sup></oasis:entry>
         <oasis:entry colname="col7">Initialization</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">PI</oasis:entry>
         <oasis:entry colname="col2">Modern</oasis:entry>
         <oasis:entry colname="col3">284.7</oasis:entry>
         <oasis:entry colname="col4">1360.9</oasis:entry>
         <oasis:entry colname="col5">Modern</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">PI_DblCO2<sup>a</sup></oasis:entry>
         <oasis:entry colname="col2">Modern</oasis:entry>
         <oasis:entry colname="col3">569.4</oasis:entry>
         <oasis:entry colname="col4">1360.9</oasis:entry>
         <oasis:entry colname="col5">Modern</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">PI</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_Ctrl</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_2x</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">569.4</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_4x</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">1138.8</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_BurlsGeo</oasis:entry>
         <oasis:entry colname="col2">
                  <xref ref-type="bibr" rid="bib1.bibx10" id="text.17"/>
                </oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1360.9</oasis:entry>
         <oasis:entry colname="col5">
                  <xref ref-type="bibr" rid="bib1.bibx10" id="text.18"/>
                </oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_noIS</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">None</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_noIS_solPI</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1360.9</oasis:entry>
         <oasis:entry colname="col5">None</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_noIS_CAM5</oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">None</oasis:entry>
         <oasis:entry colname="col6">CAM5</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_LightSoil</oasis:entry>
         <oasis:entry colname="col2">Getech &amp; higher soil albedo</oasis:entry>
         <oasis:entry colname="col3">854.1</oasis:entry>
         <oasis:entry colname="col4">1360.9</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Default</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_Ctrl_DblCO2<sup>a</sup></oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">1708.2</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Mio_Ctrl</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Mio_2x_DblCO2<sup>a</sup></oasis:entry>
         <oasis:entry colname="col2">Getech</oasis:entry>
         <oasis:entry colname="col3">1138.8</oasis:entry>
         <oasis:entry colname="col4">1359.5</oasis:entry>
         <oasis:entry colname="col5">Getech</oasis:entry>
         <oasis:entry colname="col6">CAM4</oasis:entry>
         <oasis:entry colname="col7">Mio_2x</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Methods</title>
      <p id="d2e734">We use the Community Earth System Model version 1.2 (CESM1.2) to simulate mid-Miocene climate states. CESM1.2 is a coupled model using the ocean model POP2, with a 1° horizontal grid and 60 vertical levels and the CAM4 model for the atmosphere <xref ref-type="bibr" rid="bib1.bibx45" id="paren.19"/>, applying a horizontal grid of <inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:mn mathvariant="normal">1.9</mml:mn><mml:mo>×</mml:mo><mml:mn mathvariant="normal">2.5</mml:mn><mml:mi mathvariant="italic">°</mml:mi></mml:mrow></mml:math></inline-formula> with 26 vertical layers, as well as a sea-ice model (CICE4), a river runoff model (RTM) and a land model which includes the carbon-nitrogen cycle and dynamical vegetation (CLM4). CESM1.2 has been used in numerous studies simulating different paleoclimates, such as the Last Glacial Maximum (LGM) <xref ref-type="bibr" rid="bib1.bibx34 bib1.bibx70" id="paren.20"/>, the Pliocene <xref ref-type="bibr" rid="bib1.bibx28" id="paren.21"/>, the Eocene <xref ref-type="bibr" rid="bib1.bibx39" id="paren.22"/> as well as older paleoclimates <xref ref-type="bibr" rid="bib1.bibx36" id="paren.23"/>.</p>

      <fig id="F1" specific-use="star"><label>Figure 1</label><caption><p id="d2e769">Topography and bathymetry for the two main paleogeographies in our simulations: <bold>(A)</bold> paleogeography of Mio_Ctrl, using Getech Plc. and <bold>(B)</bold> paleogeography of Mio_BurlsGeo based on <xref ref-type="bibr" rid="bib1.bibx10" id="text.24"/>. <bold>(C)</bold> is the difference in topography between both paleogeographies. Except Mio_BurlsGeo, all simulations use <bold>(A)</bold> Getech Plc.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f01.png"/>

      </fig>

      <p id="d2e793">We perform a “best-to-knowledge” control simulation of the mid-Miocene (Mio_Ctrl), which includes an Antarctic ice sheet reconstruction, an updated CO<sub>2</sub> estimate, mid-Miocene solar constant, and is based on a topography and bathymetry reconstruction of the mid-Miocene, provided by Getech Plc. This reconstruction is based on geological data which include tectonics, sediment deposit and morphology, and which is fed into a tectonic model to generate paleogeographies at <inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.1</mml:mn><mml:mi mathvariant="italic">°</mml:mi><mml:mo>×</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn><mml:mi mathvariant="italic">°</mml:mi></mml:mrow></mml:math></inline-formula> resolution, similarly as <xref ref-type="bibr" rid="bib1.bibx38" id="text.25"/>. The original geography as well as the plate model cannot be shared; however, the <inline-formula><mml:math id="M22" display="inline"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mi mathvariant="italic">°</mml:mi><mml:mo>×</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mi mathvariant="italic">°</mml:mi></mml:mrow></mml:math></inline-formula> resolution used by CESM1.2 is shown in Fig. <xref ref-type="fig" rid="F1"/> and compared to an identical simulation using the  paleogeography of MioMIP2 Phase 1 <xref ref-type="bibr" rid="bib1.bibx10" id="paren.26"/>.</p>
      <p id="d2e847">At the regional scale, the paleogeographies differ in both bathymetry and orography. The Getech paleogeography used in Mio_Ctrl features  a notably shallower Arctic basin and generally higher elevations in mountainous and plateau regions compared to the Burls paleogeography. Due to lapse-rate effects, some of the surface temperature differences between the simulations likely arise directly from these topographic differences, particularly in regions such as the Rocky Mountains, the Andes, the Tibetan Plateau, and the Antarctic ice sheet. The Getech paleogeography also includes substantial changes in the representation of the South American Pebas mega-wetlands, which are absent in the Burls paleogeography. In addition, Greenland is represented as a slightly elevated plateau in Getech, rather than hosting an ice sheet. The Getech paleogeography differs from other recent mid-Miocene paleogeographies <xref ref-type="bibr" rid="bib1.bibx18 bib1.bibx53 bib1.bibx29" id="paren.27"><named-content content-type="pre">e.g.</named-content></xref> primarily in the depths of the Fram Strait, the eastern and western seaways of the Tethys and the depth of the Panama seaway, which reflect uncertainties on the timing of closing, deepening or shoaling of these gateways during the Miocene in general. For instance, <xref ref-type="bibr" rid="bib1.bibx29" id="text.28"/> shows a much shallower eastern Tethys compared to Getech, and a western Paratethys connection, whereas <xref ref-type="bibr" rid="bib1.bibx53" id="text.29"/> displays a close eastern Tethys at the mid-Miocene. Both <xref ref-type="bibr" rid="bib1.bibx18" id="text.30"/> and <xref ref-type="bibr" rid="bib1.bibx29" id="text.31"/> show a relatively shallow Panama seaway, which differ notably from <xref ref-type="bibr" rid="bib1.bibx10" id="text.32"/>. Overall the Fram strait and earlier connections to the Arctic are much shallower in the geography of Getech than these other paleogeographies we are comparing to. To our knowledge, none of these geographies explicitly include the Pebas wetlands in South America. This may partly reflect differences in how wetlands are represented in climate models; in our case, the Pebas wetlands were treated as the shallowest level of the ocean model. Finally, paleogeographic reconstructions also differ in soil properties and vegetation type. In our simulations, vegetation type is simulated using a dynamic scheme.</p>
      <p id="d2e871">The extent of Arctic and Antarctic ice sheets between Mio_Ctrl and Mio_BurlsGeo is compared to PI in Fig. <xref ref-type="fig" rid="F2"/>. The ice sheets are considered as topographic objects and therefore the topography is flattened in Antarctica in the experiments where the ice sheets are not included.  We then compare several Miocene simulations that differ in their solar constant (Mio_noIS_solPI), the presence of Antarctic ice sheet (Mio_noIS), and the atmospheric model used (Mio_noIS_CAM5), as to identify the role of each onto the mid-Miocene climate state. A summary of each experiment is given in Table <xref ref-type="table" rid="T1"/>.</p>

      <fig id="F2" specific-use="star"><label>Figure 2</label><caption><p id="d2e880">Land ice extent (filled contours) in the Arctic (top row) and the Antarctic (bottom row) regions in the three geographies used in this study: <bold>(A)</bold> PI, <bold>(B)</bold> Mio_Ctrl with the paleogeography of Getech and <bold>(C)</bold>, Mio_BurlsGeo with the paleogeography of <xref ref-type="bibr" rid="bib1.bibx10" id="text.33"/>. Note that the PI geography also includes high altitude glaciers, which are not all shown. Land masses are in white.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f02.png"/>

      </fig>

      <p id="d2e901">Unless specified for our sensitivity tests, our simulations use an atmospheric CO<sub>2</sub> concentration of 854.1 ppm, which is equivalent to three times PI CO<sub>2</sub> concentration of the model. This is planned as the standard concentration for the protocol of the MioMIP-phase 2 project. It is within the range of some reconstructions provided for the mid-Miocene <xref ref-type="bibr" rid="bib1.bibx48 bib1.bibx58" id="paren.34"/>, and it has also been used in other mid-Miocene simulations, which in general provides a closer match to proxy temperatures than simulations with lower CO<sub>2</sub> concentrations <xref ref-type="bibr" rid="bib1.bibx10" id="paren.35"/>. The solar constant is calculated from a model of solar luminosity increase with time <xref ref-type="bibr" rid="bib1.bibx23" id="paren.36"/>, which leads to a value of 1359.5 W m<sup>−2</sup> for the mid-Miocene, in comparison to 1360.9 W m<sup>−2</sup> at PI. Aerosols are kept as PI, but the sources are moved to match with the mid-Miocene paleogeography (for instance if an ocean turned to land). Non-CO<sub>2</sub> greenhouse gases and orbital forcing are kept as PI. Our simulations are compared to a proxy record presented in <xref ref-type="bibr" rid="bib1.bibx10" id="text.37"/>. It is an agglomeration of sea-surface and surface temperatures based on multi-proxy reconstructions, where the sites are then rotated onto the mid-Miocene geography using the tectonic model of Getech Plc.</p>
      <p id="d2e977">For estimating Miocene ECS in our simulation, we base our approach on a variation of <xref ref-type="bibr" rid="bib1.bibx24" id="text.38"/>, which would require an equilibrium simulation of the mid-Miocene with PI CO<sub>2</sub> concentration, from where a new simulation is initialized with an abrupt and sustained quadrupling of the CO<sub>2</sub> concentration, for around 150 years, so that “fast” climate feedbacks (i.e. clouds, surface albedo, water vapor, lapse-rate and Planck feedbacks) have mostly responded to the initial forcing. In the case of our simulations, a mid-Miocene state with PI CO<sub>2</sub> is numerically unstable, therefore we initialize from Mio_Ctrl by instead abruptly doubling its CO<sub>2</sub> concentration (Mio_Ctrl_DblCO2), as to avoid reaching excessively high CO<sub>2</sub> concentrations under a quadrupling, which could introduce non-linear effects on climate feedbacks. The ECS is then extrapolated from the global mean annual surface temperature after 150 years of simulation minus the initial temperature of the simulation using ordinary least squares linear regression. As to roughly estimate the influence of the Miocene background temperature on ECS, we also apply the same method but from a simulation at lower CO<sub>2</sub> concentration, Mio_2x (Mio_2x_DblCO2), and we compare it to a PI simulation with an abrupt doubling of CO<sub>2</sub> (PI_DblCO2).</p>
      <p id="d2e1048">The model is initialized from a warm ocean state, where the average deep-ocean temperature is around 6.6 °C. This value comes from an earlier, unpublished mid-Miocene simulation performed with CESM1.2, and is used to accelerate our deep-ocean equilibrium. The salinity is initialized globally at 35 PSU. In South America, in the so-called Lake Pebas, salinity becomes negative due to excessive precipitation and river runoff in these warm conditions. Therefore, we apply a marginal sea balancing scheme which redistributes the gain of freshwater onto the world oceans to avoid the spread of nonphysical, negative salinity into the oceans. This scheme has previously been used with CESM1.2 in warm paleoclimates <xref ref-type="bibr" rid="bib1.bibx39" id="paren.39"><named-content content-type="pre">e.g.</named-content></xref>. The sea-ice model is initialized ice-free and the dynamic vegetation is initialized as bare soil. All runs have been integrated for at least 3100 years and up to 5000 years when using CAM5 (experiment Mio_noIS_CAM5), starting from default initial conditions, and the last 100 years are averaged and used for the results.</p>
      <p id="d2e1056">Several aspects of the ocean model are modified to handle the large changes in land-sea mask and bathymetry at the mid-Miocene. Overflows and tidal mixing are turned off, as they are hard-coded on a pre-industrial ocean grid. We follow the recommendations for deep-time paleoclimates of the National Center for Atmospheric Research (NCAR) to increase ocean background vertical diffusivity, to compensate for the lack of tidal mixing (found at <uri>https://www2.cesm.ucar.edu/models/paleo/faq/</uri>, last access: 11 April 2025). The same approach was taken by <xref ref-type="bibr" rid="bib1.bibx3" id="text.40"/> using CESM1.0.5 to simulate the Eocene. The background vertical diffusivity <inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi>w</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is horizontally homogenous but depends on model depth, and corresponds to <inline-formula><mml:math id="M37" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi>w</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mi mathvariant="normal">vdc</mml:mi><mml:mn mathvariant="normal">1</mml:mn><mml:mo>+</mml:mo><mml:mi mathvariant="normal">vdc</mml:mi><mml:mn mathvariant="normal">2</mml:mn><mml:msup><mml:mi mathvariant="normal">tan</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mo>(</mml:mo><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:mi>z</mml:mi><mml:mo>|</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="normal">depth</mml:mi><mml:mo>)</mml:mo><mml:mi mathvariant="normal">linv</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where vdc1 <inline-formula><mml:math id="M38" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.524</mml:mn></mml:mrow></mml:math></inline-formula> cm<sup>2</sup> s<sup>−1</sup>, vdc2 <inline-formula><mml:math id="M41" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.313</mml:mn></mml:mrow></mml:math></inline-formula> cm<sup>2</sup> s<sup>−1</sup>, depth <inline-formula><mml:math id="M44" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1000</mml:mn></mml:mrow></mml:math></inline-formula> m, and linv <inline-formula><mml:math id="M45" display="inline"><mml:mrow><mml:mo>=</mml:mo><mml:mn mathvariant="normal">4.5</mml:mn><mml:mo>×</mml:mo><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mrow><mml:mi mathvariant="normal">−</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> m<sup>−1</sup>. This equation of background vertical diffusivity follows a <xref ref-type="bibr" rid="bib1.bibx7" id="text.41"/> tangential profile, where the upper ocean diffusivity is 0.1 cm<sup>2</sup> s<sup>−1</sup> and the deep ocean diffusivity is a constant 1 cm<sup>2</sup> s<sup>−1</sup> below 1000 m. As a comparison, the PI state overall has a lower background diffusivity in the deep ocean, but the bottom-intensified tidal mixing can be up to two orders of magnitude larger than the background mixing. The timestep of the ocean model is reduced from calculating its step every 63 min model-time, to 41 min model-time. Modifying the timestep of models which use numerical integrations is known to affect their results, as was shown for CAM3 <xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx67" id="paren.42"/>, yet it remains generally poorly documented. It is not uncommon for paleoclimate simulations to use shorter computational time steps, as the ocean model in particular can generate numerous numerical instabilities which can lead to an early crash of the model. Our mid-Miocene results are compared to the “best-to-knowledge” PI simulation using modern parameterizations, as this should more accurately resemble the PI climate state. we explore the impact of these choice of ocean parametrizations in Appendix <xref ref-type="sec" rid="App1.Ch1.S1"/>, as further work on the ocean component of our mid-Miocene simulations is kept for a future study.</p>

      <fig id="F3" specific-use="star"><label>Figure 3</label><caption><p id="d2e1287">Surface temperature <bold>(A)</bold> and precipitation <bold>(D)</bold> maps of Mio_Ctrl, as well as comparison with PI <bold>(B, E)</bold> and with Mio_BurlsGeo <bold>(C, F)</bold>.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f03.png"/>

      </fig>

</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Mid-Miocene control state</title>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Climate</title>
      <p id="d2e1323">Our “best-to-knowledge” mid-Miocene simulation (Mio_Ctrl), which includes changes in paleogeography, ice sheet, CO<sub>2</sub> concentration, solar forcing and vegetation, is compared to PI and a similar mid-Miocene simulation using the paleogeography provided for MioMIP2 Phase 1 (Mio_BurlsGeo), in Fig. <xref ref-type="fig" rid="F3"/>. Mio_Ctrl simulates a GMST of 19.8 °C, which is 5.4 °C warmer than PI (Table <xref ref-type="table" rid="T2"/> and Fig. <xref ref-type="fig" rid="F3"/>A). This temperature anomaly is in line with other simulations of the mid-Miocene Climate Optimum (MMCO) at lower CO<sub>2</sub> concentrations <xref ref-type="bibr" rid="bib1.bibx31 bib1.bibx66 bib1.bibx61" id="paren.43"/>, and is around 2 °C colder than the mid-Miocene simulations performed with HadCM3L at similar CO<sub>2</sub> concentrations <xref ref-type="bibr" rid="bib1.bibx10" id="paren.44"/>.</p>
      <p id="d2e1366">Regionally, our mid-Miocene simulations (Mio_Ctrl and Mio_BurlsGeo) both show a substantial warming over the southern high latitudes (Fig. <xref ref-type="fig" rid="F3"/>B), linked to polar amplification, and a large cooling pattern over the north Atlantic ocean and northern Europe. The latter is likely connected to an absent Atlantic Meridional Overturning Circulation (AMOC), which is replaced by a Pacific Meridional Overturning Circulation (PMOC). The absent AMOC implies a reduced northward heat transport in the North Atlantic and towards the Arctic ocean. The Arctic, in turn, is shallow and has limited exchanges with the Atlantic ocean in these paleogeographies. This likely contributes to the apparent absence of polar amplification in the northern hemisphere temperature signal. An absent AMOC in favor of the PMOC has been seen in other mid-Miocene simulations but remains a rare phenomenon within MioMIP, and is not supported by proxy data <xref ref-type="bibr" rid="bib1.bibx32 bib1.bibx44" id="paren.45"/>. Although this result is relevant for future climate change and for understanding the conditions under which a PMOC state can be sustained, it will be addressed in a separate paper focusing on the ocean component of our simulations.</p>
      <p id="d2e1374">Comparing both paleogeographies, Mio_BurlsGeo is slightly warmer than Mio_Ctrl by 0.2 °C (Table <xref ref-type="table" rid="T2"/> and Fig. <xref ref-type="fig" rid="F3"/>C). Mio_BurlsGeo was initialized with a slightly lighter color of soil, equivalent to a dry albedo increase of around 0.1 and a saturated soil albedo increase of around 0.05 compared to the darker soil of Mio_Ctrl and other simulations using the Getech paleogeography. Soil properties are rarely documented in past modelling studies, mainly due to the fact that there is limited information on soil albedo in geological times. However, testing the effect of soil properties can be relevant in future paleomodelling efforts, in particular as it interacts with dynamic vegetation. Adjusting the soil color of Mio_Ctrl to that of Mio_BurlsGeo in a simulation called Mio_LightSoil leads to a GMST of around 18.7 °C, and so a difference with Mio_BurlsGeo of around 1.3 °C. While there is limited constraint on soil properties in paleo settings, and it is largely dependent of i.e. soil type, humidity, vegetation cover, ice sheet, we emphasize that documenting these properties is relevant as they may partially contribute to the mid-Miocene climatic patterns.</p>
      <p id="d2e1381">Mio_BurlsGeo was also initialized with PI solar forcing as to provide a better comparison with previously published mid-Miocene runs. As discussed in Sect. <xref ref-type="sec" rid="Ch1.S6.SS3"/>, a simulation with PI solar forcing (Mio_noIS_solPI) shows an additional warming of around 1 °C in high latitudes. However, as also discussed in Sect. <xref ref-type="sec" rid="Ch1.S6.SS3"/>, the difference of solar constant between Mio_BurlsGeo and Mio_Ctrl is mostly negligible compared to the other forcing discussed in this study.</p>
      <p id="d2e1389">Similarly as for temperature, we analyse total precipitation in Mio_Ctrl (Fig. <xref ref-type="fig" rid="F3"/>D) with respect to PI and Mio_BurlsGeo. Globally, Mio_Ctrl is 12.9 % wetter than PI (Table <xref ref-type="table" rid="T2"/> and Fig. <xref ref-type="fig" rid="F3"/>E) and is wetter than mid-Miocene simulations using HadCM3L at 850 ppm of CO<sub>2</sub> (around 10 %, <xref ref-type="bibr" rid="bib1.bibx1" id="altparen.46"/>). Mio_Ctrl is characterized by drier conditions in the eastern Pacific and wetter conditions in the western Pacific, as well as a common double Intertropical Convergence Zone (ITCZ) bias over the basin <xref ref-type="bibr" rid="bib1.bibx57" id="paren.47"/>. This is in good agreement with the multi-model mean of mid- to late Miocene simulations at 560 ppm of CO<sub>2</sub> <xref ref-type="bibr" rid="bib1.bibx1" id="paren.48"/>. The north Atlantic region is drier than PI, and similar results are obtained from HadCM3L and CCSM4 simulations <xref ref-type="bibr" rid="bib1.bibx10 bib1.bibx1" id="paren.49"/>.</p>
      <p id="d2e1429">Mio_BurlsGeo is slightly wetter than Mio_Ctrl (Fig. <xref ref-type="fig" rid="F3"/>F), and 14 % wetter than PI (Table <xref ref-type="table" rid="T2"/>). Precipitation patterns are partly affected by the differences in coastlines and topography as seen in Fig. <xref ref-type="fig" rid="F1"/>. In the tropics, the absence of the Pebas wetlands in Mio_BurlsGeo results in drier conditions in the northern part of South America. In contrast, the South-East Pacific, North-East Asia and South-East Asia regions are wetter and the latter is, in fact, the largest precipitation change between both geographies. The main characteristic of this region is an enhanced connection between the Pacific and the Indian oceans which is not present in Mio_Ctrl.</p>

<table-wrap id="T2" specific-use="star"><label>Table 2</label><caption><p id="d2e1441">Summary of global mean surface temperature (GMST) and the anomaly relative to the PI simulation, and percent change of total precipitation (%<inline-formula><mml:math id="M56" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>p) relative to PI in our simulations.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="center"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">#</oasis:entry>
         <oasis:entry colname="col2">Run</oasis:entry>
         <oasis:entry colname="col3">GMST (°C)</oasis:entry>
         <oasis:entry colname="col4">GMST anomaly (°C)</oasis:entry>
         <oasis:entry colname="col5">%<inline-formula><mml:math id="M57" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>p</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">0</oasis:entry>
         <oasis:entry colname="col2">PI</oasis:entry>
         <oasis:entry colname="col3">14.4</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">1</oasis:entry>
         <oasis:entry colname="col2">Mio_Ctrl</oasis:entry>
         <oasis:entry colname="col3">19.8</oasis:entry>
         <oasis:entry colname="col4">5.4</oasis:entry>
         <oasis:entry colname="col5">12.9</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">2</oasis:entry>
         <oasis:entry colname="col2">Mio_2x</oasis:entry>
         <oasis:entry colname="col3">18.0</oasis:entry>
         <oasis:entry colname="col4">3.6</oasis:entry>
         <oasis:entry colname="col5">9.4</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">3</oasis:entry>
         <oasis:entry colname="col2">Mio_4x</oasis:entry>
         <oasis:entry colname="col3">21.2</oasis:entry>
         <oasis:entry colname="col4">6.8</oasis:entry>
         <oasis:entry colname="col5">15.3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">4</oasis:entry>
         <oasis:entry colname="col2">Mio_noIS</oasis:entry>
         <oasis:entry colname="col3">18.5</oasis:entry>
         <oasis:entry colname="col4">4.1</oasis:entry>
         <oasis:entry colname="col5">9.0</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">5</oasis:entry>
         <oasis:entry colname="col2">Mio_noIS_solPI</oasis:entry>
         <oasis:entry colname="col3">18.7</oasis:entry>
         <oasis:entry colname="col4">4.3</oasis:entry>
         <oasis:entry colname="col5">9.5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">6</oasis:entry>
         <oasis:entry colname="col2">Mio_BurlsGeo</oasis:entry>
         <oasis:entry colname="col3">20.0</oasis:entry>
         <oasis:entry colname="col4">5.6</oasis:entry>
         <oasis:entry colname="col5">14.0</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">7</oasis:entry>
         <oasis:entry colname="col2">Mio_noIS_CAM5</oasis:entry>
         <oasis:entry colname="col3">21.7</oasis:entry>
         <oasis:entry colname="col4">7.3</oasis:entry>
         <oasis:entry colname="col5">20.0</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">8</oasis:entry>
         <oasis:entry colname="col2">Mio_LightSoil</oasis:entry>
         <oasis:entry colname="col3">18.7</oasis:entry>
         <oasis:entry colname="col4">4.3</oasis:entry>
         <oasis:entry colname="col5">10</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">9</oasis:entry>
         <oasis:entry colname="col2">PI_DblCO2</oasis:entry>
         <oasis:entry colname="col3">17.2</oasis:entry>
         <oasis:entry colname="col4">2.8</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">10</oasis:entry>
         <oasis:entry colname="col2">Mio_Ctrl_DblCO2</oasis:entry>
         <oasis:entry colname="col3">22.7</oasis:entry>
         <oasis:entry colname="col4">8.3</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">11</oasis:entry>
         <oasis:entry colname="col2">Mio_2x_DblCO2</oasis:entry>
         <oasis:entry colname="col3">20.6</oasis:entry>
         <oasis:entry colname="col4">6.2</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Vegetation</title>
      <p id="d2e1724">Biomes based on vegetation distribution are challenging to reconstruct in deep time paleoclimates, notably due to the difficulties in preserving land proxy data and because the plant pollen can be deposited far from their sources. However, vegetation plays an important  role in past climates as it provides feedbacks on moisture and temperature changes, and can be used to reconstruct atmospheric CO<sub>2</sub> concentrations. The mid-Miocene is particularly relevant as it displays the largest flora change of the Cenozoic and the emergence of modern biomes <xref ref-type="bibr" rid="bib1.bibx58" id="paren.50"/>.</p>
      <p id="d2e1739">Coupled climate-vegetation models have been used in past climate simulations <xref ref-type="bibr" rid="bib1.bibx60" id="paren.51"><named-content content-type="pre">e.g.</named-content></xref>, with relatively good agreement with proxy data, albeit large local differences can be seen. Here, we show the simulated vegetation for Mio_Ctrl (Fig. <xref ref-type="fig" rid="F4"/>) and compare it to the reconstructed biomes for the mid-Miocene of <xref ref-type="bibr" rid="bib1.bibx58" id="text.52"/>.</p>

      <fig id="F4" specific-use="star"><label>Figure 4</label><caption><p id="d2e1754"><bold>(A)</bold> Simulated vegetation types in Mio_Ctrl compared to <bold>(B)</bold> reconstructed biomes for the mid-Miocene, adapted from <xref ref-type="bibr" rid="bib1.bibx58" id="text.53"/>. In <bold>(A)</bold>, the soil is bare if the sum of the contribution of all vegetated areas is less than 10 %.</p></caption>
          <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f04.png"/>

        </fig>

      <p id="d2e1775">The proxy record indicates a large spread of temperate forests and savanna in mid-latitudes during the mid-Miocene, notably in Europe, north-east Asia and Australia. Although our vegetation model also simulates temperate biomes for Europe and Australia, predominantly composed of temperate shrubs or evergreen temperate forests, we observe Arctic (dominated by Arctic grass) and boreal (dominated by boreal evergreen trees) climates in North East Asia and northern Europe, and almost no vegetation growth above 60° N. On the contrary, the proxy record still indicates (cool) temperate forests, even at latitudes higher than 60° N.</p>
      <p id="d2e1778">Overall, our predicted vegetation mirrors our model-data temperature map: there is a good agreement in tropical regions between model and proxy data, but our simulated biomes are contracted towards the Equator, with warmer types of vegetation constrained in the lower latitudes and therefore fail to match mid to high latitude biomes as seen in the proxy record. This is likely both a result of and feedback on temperature, as vegetation usually has a lower albedo than bare soil.</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Impact of CO<sub>2</sub></title>
      <p id="d2e1799">There have been different plausible estimates of atmospheric CO<sub>2</sub> concentrations for the mid-Miocene due to differences in methodology and calibrations <xref ref-type="bibr" rid="bib1.bibx10" id="paren.54"/>. The concentration is usually thought to lie between two and three times PI concentrations, three being the concentration discussed for the upcoming MioMIP experimental design, in line with recent high estimates during the MMCO (around 700–1100 ppm, <xref ref-type="bibr" rid="bib1.bibx48" id="altparen.55"/>). Here, we compare Mio_Ctrl, which has a CO<sub>2</sub> concentration of three times PI, to two simulations using mid-Miocene boundary conditions and two times (Mio_2x) and four times (Mio_4x) PI in Fig. <xref ref-type="fig" rid="F5"/>, relative to the proxy record. Although CO<sub>2</sub> concentrations as high as four times PI during the mid-Miocene are at the high threshold of the estimates, such levels are useful for estimating ECS and feedbacks <xref ref-type="bibr" rid="bib1.bibx16" id="paren.56"/>, which are both important elements for quantifying global and regional responses to CO<sub>2</sub>.</p>
      <p id="d2e1850">In Mio_2x, the GMST is 18.0 °C, which is 3.6 °C warmer than PI (Table <xref ref-type="table" rid="T2"/>), and even lower than proxy estimates of the colder Late Miocene <xref ref-type="bibr" rid="bib1.bibx10" id="paren.57"/>. In Mio_4x, the GMST is 21.2 °C, and its anomaly relative to PI is 6.8 °C (Table <xref ref-type="table" rid="T2"/>). This is in agreement with proxy estimates for the GMST of the mid-Miocene. Similarly, <xref ref-type="bibr" rid="bib1.bibx1" id="text.58"/> showed that CO<sub>2</sub> concentrations between three and four times PI concentrations provide the most reduced bias in total precipitation at a global scale, but some regions critically lack proxy sites, and there are few simulations at such high CO<sub>2</sub> concentrations for comparison <xref ref-type="bibr" rid="bib1.bibx10" id="paren.59"/>.</p>
      <p id="d2e1885">At all CO<sub>2</sub> levels, the mid-Miocene simulations are cold and underestimate surface temperatures reconstructed from proxy data in most mid to high latitudes locations (Fig. <xref ref-type="fig" rid="F5"/>). The cold pattern simulated in the north Atlantic contributes even more to the temperature bias relative to proxy data. This pattern is likely connected to an absent AMOC in our model at all three CO<sub>2</sub> levels. Problematically, a weak mid-Miocene AMOC is a persistent feature in climate models <xref ref-type="bibr" rid="bib1.bibx44" id="paren.60"/>, with some rare exceptions <xref ref-type="bibr" rid="bib1.bibx62" id="paren.61"/>, the associated colder North Atlantic surface temperature is not necessarily supported by proxies <xref ref-type="bibr" rid="bib1.bibx54" id="paren.62"><named-content content-type="pre">e.g.</named-content></xref>. Nevertheless, a recent study using coccolith clumped isotopes showed a modest northern Atlantic warming during the mid-Miocene compared to the high warming reported in earlier studies <xref ref-type="bibr" rid="bib1.bibx41" id="paren.63"/>, which suggest that some model-proxy discrepancies may be due to limitations in proxy interpretation. Further discussion on the ocean circulation in our simulations is planned for a follow-up paper.</p>
      <p id="d2e1924">In Mio_2x, the temperatures are too cold even in tropical locations compared to proxy data (Fig. <xref ref-type="fig" rid="F5"/>). On the contrary, in Mio_4x, some of the mid and high latitudes negative biases are dampened, but tropical locations are too warm compared to proxy data. These issues affecting polar and tropical warmth are commonly found in other Miocene simulations with various models <xref ref-type="bibr" rid="bib1.bibx10" id="paren.64"/>, and most mid-Pliocene simulations during PlioMIP1 <xref ref-type="bibr" rid="bib1.bibx27" id="paren.65"/>, although there was no systematic bias in PlioMIP2 models <xref ref-type="bibr" rid="bib1.bibx28" id="paren.66"/>. Climate models are not tuned for paleoclimates, which contributes to the challenges they face in representing polar amplified temperatures in warm paleoclimates <xref ref-type="bibr" rid="bib1.bibx9" id="paren.67"/>. Moreover, climate models may show other biases in high latitudes, notably due to poor representation of vegetation from dynamic vegetation models, as shown in Sect. <xref ref-type="sec" rid="Ch1.S3.SS2"/>, or lack of interactive ice sheets. Additionally, it is likely that model-proxy biases are a combination of both model and proxy errors. Notably, pollen and marine organic proxies can have substantial seasonal biases, which can strongly affect their temperature signal when compared to annually-averaged modelled temperatures, as described by <xref ref-type="bibr" rid="bib1.bibx64" id="text.68"/> for the Pliocene.</p>

      <fig id="F5" specific-use="star"><label>Figure 5</label><caption><p id="d2e1949">Surface temperature biases relative to the proxy record at <bold>(A)</bold> 2 (Mio_2x), <bold>(B)</bold> 4 (Mio_4x) and <bold>(C)</bold> 3 (Mio_Ctrl) times PI CO<sub>2</sub> concentrations, at proxy sites and <bold>(D)</bold> zonally. Proxy sites are rotated on the mid-Miocene geography with the tectonic model of Getech Plc.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f05.png"/>

      </fig>

</sec>
<sec id="Ch1.S5">
  <label>5</label><title>Equilibrium Climate Sensitivity</title>
      <p id="d2e1988">ECS is widely regarded as one of the most critical metrics for characterizing future climate change <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx33" id="paren.69"/>. However, due to the difficulty of constraining it, several independent methods have been used, notably including evidence from the paleoclimate record <xref ref-type="bibr" rid="bib1.bibx15 bib1.bibx26 bib1.bibx52 bib1.bibx50 bib1.bibx55" id="paren.70"><named-content content-type="pre">e.g.</named-content></xref>. Some of these methods involve directly calculating the radiative forcing and global temperature of a past climate from the proxy record <xref ref-type="bibr" rid="bib1.bibx52 bib1.bibx55 bib1.bibx63" id="paren.71"><named-content content-type="pre">e.g</named-content></xref>, or emergent-constraint approaches, which are statistical methods connecting proxy reconstruction with the outputs from several models, usually under the umbrella of a MIP, such as for the LGM under PMIP <xref ref-type="bibr" rid="bib1.bibx26 bib1.bibx50 bib1.bibx51" id="paren.72"/> or PlioMIP <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx28 bib1.bibx51" id="paren.73"/>.</p>
      <p id="d2e2010">The ECS of a paleoclimate and how it relates to modern ECS is not necessarily obvious. The effects of “fast” climate feedbacks (i.e. clouds, surface albedo, water vapor, lapse-rate and Planck feedbacks) need to be disentangled from “slow” climate feedbacks (notably carbon cycle, ice sheets and aerosols feedbacks), where the latter affects temperature on geological time-scale and usually connect to a different concept, the Earth system sensitivity <xref ref-type="bibr" rid="bib1.bibx52 bib1.bibx37" id="paren.74"/>. Climate feedbacks also depend on the background climate and the forcing imposed, which can lead to high ECS in warm, high-CO<sub>2</sub> paleoclimates <xref ref-type="bibr" rid="bib1.bibx12 bib1.bibx2" id="paren.75"/>. Those aspects result in important challenges when using evidence from new paleo-MIP to deduce ECS <xref ref-type="bibr" rid="bib1.bibx52 bib1.bibx51" id="paren.76"/>.</p>
      <p id="d2e2031">A way to approach these issues is to estimate ECS directly from a paleoclimate simulation, and compare it to the ECS in a modern simulation in the same model <xref ref-type="bibr" rid="bib1.bibx16" id="paren.77"><named-content content-type="pre">e.g.</named-content></xref>. Here, we have performed two simulations: Mio_Ctrl_DblCO2, which corresponds to a doubling of CO<sub>2</sub> from Mio_Ctrl (1708.2 ppm), and Mio_2x_DblCO2, which is a doubling of CO<sub>2</sub> from Mio_2x (1138.8 ppm). Further details on the methodology are provided in Sect. <xref ref-type="sec" rid="Ch1.S2"/>.</p>

      <fig id="F6" specific-use="star"><label>Figure 6</label><caption><p id="d2e2062">Top-of-atmosphere energy flux (W m<sup>−2</sup> versus global mean surface temperature where <bold>(A)</bold> the simulations temperature are reported relative to the PI temperature and <bold>(B)</bold> the simulation temperatures are reported relative to their initial state (PI or mid-Miocene). The stars (one for each simulation/color) are estimates of the long-term near-equilibrium temperatures using ordinary least squares linear regression.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f06.png"/>

      </fig>

      <p id="d2e2089">In Mio_Ctrl_DblCO2 (1708.2 ppm), ECS is 2.9 °C (2.5–3.3 °C, 95 % predicted interval, Fig. <xref ref-type="fig" rid="F6"/>B), in comparison to 2.8 °C from PI conditions with this model (2.2–3.4 °C, 95 % predicted interval, Fig. <xref ref-type="fig" rid="F6"/>A). From a lower CO<sub>2</sub> concentration, Mio_2x_DblCO2 (1138.8 ppm), ECS is 2.6 °C, which, similarly as the estimate from PI climate, is also within the 95 % predicted interval for the ECS of Mio_Ctrl_DblCO2. Even when considering the higher background CO<sub>2</sub> concentration, and consequently higher background mean temperature of the mid-Miocene, these estimates tend to indicate that the PI-based ECS of CESM1.2 can characterize the Miocene climate, or that ECS estimates obtained from the Miocene climate using CESM1.2, for instance in an emergent constraint approach, could be representative of its modern-based ECS. Previous studies on the Miocene have also estimated its ECS to be close to 3 °C, and in general relatively close to the ECS estimated for modern climate from Miocene evidence <xref ref-type="bibr" rid="bib1.bibx65 bib1.bibx52 bib1.bibx5 bib1.bibx6" id="paren.78"/>, as well as the ECS reported from multiple lines of evidence in the latest assessment report of the International Panel on Climate Change (IPCC AR6: 3 °C, 2–5 °C, 90 % confidence interval, <xref ref-type="bibr" rid="bib1.bibx17" id="altparen.79"/>), in comparison to some older warm paleoclimates (e.g. the Eocene) where ECS estimates easily exceed 6 °C <xref ref-type="bibr" rid="bib1.bibx2" id="paren.80"/>.</p>
</sec>
<sec id="Ch1.S6">
  <label>6</label><title>Additional sensitivity studies</title>
      <p id="d2e2132">Here, we investigate the impact of the Antarctic ice sheet, atmospheric model, and solar constant on the mid-Miocene temperature and precipitation. The sensitivity runs for the atmospheric model and the solar constant were performed without the Antarctic ice sheet, as the ice sheet data was not available to us at the time of the simulations, and these simulations are computationally expensive to perform. Therefore, in this section we use Mio_noIS, a mid-Miocene simulation without Antarctic ice sheet as the control simulation around which the sensitivities are tested. Global temperature anomalies and percent change of total precipitation relative to PI are reported in Table <xref ref-type="table" rid="T2"/>.</p>

      <fig id="F7" specific-use="star"><label>Figure 7</label><caption><p id="d2e2139">Surface temperature maps for the different sensitivity experiments, using the no-ice-sheet (Mio_noIS, panel <bold>A</bold>) experiment as a control state: <bold>(B)</bold> anomaly from Mio_noIS_CAM5, <bold>(C)</bold> anomaly from Mio_Ctrl and <bold>(D)</bold> anonaly from Mio_noIS_solPI. Note the differences in scale.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f07.png"/>

      </fig>

      <fig id="F8" specific-use="star"><label>Figure 8</label><caption><p id="d2e2162">Total precipitation maps for the different sensitivity experiments, using the no-ice-sheet (Mio_noIS, panel <bold>A</bold>) experiment as a control state: <bold>(B)</bold> anomaly from Mio_noIS_CAM5, <bold>(C)</bold> anomaly from Mio_Ctrl and <bold>(D)</bold> anonaly from Mio_noIS_solPI. Note the differences in scale.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f08.png"/>

      </fig>

<sec id="Ch1.S6.SS1">
  <label>6.1</label><title>Impact of atmospheric model</title>
      <p id="d2e2191">Model parametrizations play an important role on climate. In the Pliocene Modelling Intercomparison Project Phase 2 (PlioMIP2), three versions of CCSM4 simulated the mid-Pliocene (CCSM4-Utrecht, CCSM4-NCAR and CCSM4-UoT). The global mean surface temperature difference between the warmest CCSM4 (CCSM4-Utrecht) and the coldest CCSM4 (CCSM4-UoT) was as large as the global mean surface temperature difference between the coldest CCSM4 version and PI climate <xref ref-type="bibr" rid="bib1.bibx28" id="paren.81"/>. CCSM4 is a model of the CESM model family (notably containing CCSM3, CCSM4, CESM1.2, CESM1.3, CESM2, i.e. built on similar versions of submodel components developped at NCAR), such as CESM1.2 which we are using for our mid-Miocene simulations. Throughout the model family history, there have been substantial changes in parametrizations, with the most recent CESM2 having one of the largest ECS of modern climate model ensembles <xref ref-type="bibr" rid="bib1.bibx21" id="paren.82"/>.</p>

      <fig id="F9"><label>Figure 9</label><caption><p id="d2e2202">Map of surface temperature differences between Mio_noIS_CAM5 and PI. Note that the color mapping is non-linear.</p></caption>
          <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f09.png"/>

        </fig>

      <fig id="F10"><label>Figure 10</label><caption><p id="d2e2213">Global temperature change (°C) versus global precipitation change (%) relative to PI across all our mid-Miocene simulations (as numbered in Table <xref ref-type="table" rid="T2"/>) initialized from default model conditions.</p></caption>
          <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f10.png"/>

        </fig>

      <p id="d2e2225">Similarly as for temperature, changing the atmospheric model from CAM4 to CAM5 has a considerable effect on precipitation. Globally, we observe an increase to total precipitation of 20.0 % relative to PI, which is also 11 % larger than Mio_noIS (Table <xref ref-type="table" rid="T2"/>). Overall, there is a tight coupling between the global mean temperatures and global precipitation changes across our different simulations and independently of the choice of atmospheric model, as shown in Fig. <xref ref-type="fig" rid="F10"/>. This is to be expected from the Clausius-Clapeyron relation, and it is an indicator of the dominant role of global temperature change on global precipitation, whereas the different sensitivities tested in this study (e.g. topography, atmospheric model), are more likely to affect regional precipitation. For instance, as earlier mentioned, CAM5 saw substantial changes in mid to high latitudes cloud parametrizations and microphysics scheme relative to CAM4, where we observe much wetter conditions in those regions with CAM5 in our simulations (Fig. <xref ref-type="fig" rid="F8"/>), particulary in the Southern hemisphere and the northern Pacific ocean. This is at first not expected because CAM5 saw reduced non-convective precipitation relative to CAM4 <xref ref-type="bibr" rid="bib1.bibx13" id="paren.83"/>, so the increase in high latitude precipitation is more likely a response of CAM5 to CO<sub>2</sub> rather than differences in parametrizations between both atmospheric models. On the contrary, CAM5 displays enhanced convective precipitation relative to CAM4 <xref ref-type="bibr" rid="bib1.bibx13" id="paren.84"/>, and over the equatorial Pacific ocean, a wetter double ITCZ band is observed in future scenarios using CAM5 compared to CAM4 <xref ref-type="bibr" rid="bib1.bibx40" id="paren.85"/>, which could contribute to the substantial wettening of the equatorial Pacific ocean in Mio_noIS_CAM5. Drier conditions are observed over North America and in the eastern subtropical Pacific ocean. Contrarily to simulations using CAM4, the Pebas wetlands has a less pronounced impact on South American precipitation.</p>
      <p id="d2e2253">An issue in paleo MIPs is that they often include several versions of the same models with apparent small differences between them due to the cost and technical difficulties which surround paleoclimate modelling. This was the case with three versions of CCSM4 at PlioMIP2 <xref ref-type="bibr" rid="bib1.bibx28" id="paren.86"/> or multiple versions of CCSM3 and HadCM3L for MioMIP <xref ref-type="bibr" rid="bib1.bibx10" id="paren.87"/>. This can impact multi-model comparisons as well as model-proxy comparisons, as similar biases are replicated and have a bigger weight on the ensemble. Here, we have shown that moving from CAM4 onto CAM5 leads to a substantial difference in mid-Miocene climate. Thus, small changes of parametrizations between relatively similar models can generate a large array of responses and can easily justify the presence of these models together within a MIP.</p>
</sec>
<sec id="Ch1.S6.SS2">
  <label>6.2</label><title>Impact of Antarctic ice sheet</title>
      <p id="d2e2270">Geological evidences show a substantial retreat of the Antarctic ice sheet during the early to mid-Miocene, in particular during the MMCO <xref ref-type="bibr" rid="bib1.bibx20" id="paren.88"/>. In the two paleogeographies used in this study, we report a reduction in Southern hemisphere land ice area (which also include Andean glaciers) of 18 % using the paleogeography of Getech, and 31 % using the paleogeography of <xref ref-type="bibr" rid="bib1.bibx10" id="text.89"/> relative to our PI state. For the Northern hemisphere, there is no land ice in the Getech paleogeography, and a reduction of 80 % using the paleogeography of <xref ref-type="bibr" rid="bib1.bibx10" id="text.90"/>. The extent of the land ice is shown in Fig. <xref ref-type="fig" rid="F2"/>.</p>
      <p id="d2e2284">Through modification of the surface albedo, the topography and the atmospheric and ocean circulations, ice sheets have a substantial impact on the climate <xref ref-type="bibr" rid="bib1.bibx20" id="paren.91"/>. Here, we test the impact of the Antarctic ice sheet on the mid-Miocene climate by comparing to Mio_Ctrl a simulation where we removed the ice sheet and jointly flatten the Antarctic continent (Mio_noIS). Although there is a net reduction in the land ice area in our mid-Miocene simulations compared to PI, we would still expect the presence of ice sheets in Mio_Ctrl to cool down the climate relatively to Mio_noIS, notably through the increase of surface albedo. Instead, implementing the Antarctic ice sheet in Mio_Ctrl warms up the global climate by 1.4 °C and makes it 3.9 % wetter compared to Mio_noIS (Table <xref ref-type="table" rid="T2"/>).</p>
      <p id="d2e2292">When comparing both cases with and without ice sheet in Fig. <xref ref-type="fig" rid="F7"/>, we observe that the presence of the ice sheet locally cools down Antarctica by around 25 °C at its highest altitude. However, there is also a substantial warming in the Arctic ocean and over the northern hemisphere continental masses, as well as a smaller warming over the global ocean. Mio_noIS shows almost no warming at all in the Arctic region relative to PI, which differs from Mio_Ctrl. Over the oceans, mid and low latitude regions are drier or wetter in Mio_Ctrl if they are respectively dry and wet in Mio_noIS. High latitudes tend to be wetter in Mio_Ctrl, except directly over the Antarctic ice sheet. A substantial wettening of Eastern Africa can be observed in Mio_Ctrl, as well as Southern Asia.</p>
      <p id="d2e2297">Atmospheric and oceanic heat transport are similar between the simulations, indicating that the higher Northern Hemisphere temperatures are not due to enhanced heat transport from the south (Appendix <xref ref-type="sec" rid="App1.Ch1.S2"/>). Instead, the radiation budget in the northern high latitudes suggests that the warming could be a result of a complex interplay between cloud and ice fraction forces and feedbacks. A detailed investigation of these mechanisms is beyond the scope of this study, but we briefly discussed it in Appendix <xref ref-type="sec" rid="App1.Ch1.S2"/>, as it could highlight some processes which affect polar amplification in warm paleo-simulations.</p>
</sec>
<sec id="Ch1.S6.SS3">
  <label>6.3</label><title>Impact of solar constant</title>
      <p id="d2e2312">Of all the forcing applied to our mid-Miocene simulations, the change from PI solar constant to mid-Miocene solar constant has the least impact. Mio_noIS_solPI is around 0.2 °C warmer than Mio_noIS (Table <xref ref-type="table" rid="T2"/>) globally, whereas most of the warming comes from high latitudes where the temperatures are 1 °C warmer (Fig. <xref ref-type="fig" rid="F7"/>). The small amplitude of this change is expected, as the mid-Miocene solar constant is estimated to be 0.1 % less than PI solar constant <xref ref-type="bibr" rid="bib1.bibx23" id="paren.92"/>. As a comparison, a doubling of atmospheric CO<sub>2</sub> (around 3.7 W m<sup>−2</sup>, <xref ref-type="bibr" rid="bib1.bibx43" id="altparen.93"/>) is roughly equivalent in forcing to an increase of 2.25 % of solar constant, assuming a planetary albedo of 0.7. Therefore, the change of solar constant is mostly negligible, but could be partially responsible for the high latitudes of Mio_BurlsGeo being warmer than Mio_Ctrl which uses a mid-Miocene solar constant.</p>
</sec>
</sec>
<sec id="Ch1.S7" sec-type="conclusions">
  <label>7</label><title>Conclusions</title>
      <p id="d2e2356">In this study, we have developed a set of simulations using a new and previously unpublished geography for the mid-Miocene, using CESM1.2. Research centered on the mid-Miocene has shown a potential for it to be a partial analog of near-future climate <xref ref-type="bibr" rid="bib1.bibx58" id="paren.94"><named-content content-type="pre">e.g.</named-content></xref>, yet uncertainties remain on atmospheric CO<sub>2</sub> concentration and non-CO<sub>2</sub> forcing, and consequently the scale of the warming and the strength of its potential analogy. In particular, model simulations do not match the high latitude warming inferred from some proxies, potentially due to changes in ocean circulation, poor understanding of polar amplification in models or poor representation of vegetation. Alternatively, proxy-reconstructed temperatures might also be uncertain, as recent studies suggest colder mid-Miocene northern Atlantic temperatures, more in-line with climate models' outputs <xref ref-type="bibr" rid="bib1.bibx41" id="paren.95"/>. It is in fact likely that these discrepancies reflect issues from both models and proxies, and the mid-Miocene experiments performed in this study aimed to bring a new light onto those questions.</p>
      <p id="d2e2385">Variations in CO<sub>2</sub> concentrations for the mid-Miocene have a substantial influence on its estimated temperatures. Between the high end (four times PI concentration) and the low end (two times PI concentration) of tested concentrations, we find a difference of 3.2 °C on mid-Miocene global mean surface temperature, suggesting a substantial role of CO<sub>2</sub> forcing on the mid-Miocene climate, as also previously observed in the case of the Pliocene <xref ref-type="bibr" rid="bib1.bibx11" id="paren.96"/>. The mid-Miocene is also sensitive to non-CO<sub>2</sub> forcing, such as the land-sea mask, ice sheet and solar forcing, but the variations tested for those components had a lower influence on the mid-Miocene temperature than CO<sub>2</sub>. Finally, we identified that switching the atmospheric model from CAM4 to CAM5, which has a substantial influence on cloud feedbacks, leads to a warmer mid-Miocene by up to 3.2 °C, which highlights the critical need of improving model climate feedbacks, as much as it is to better constrain climate forcing in paleoclimates.</p>
      <p id="d2e2427">Similarly as for the LGM <xref ref-type="bibr" rid="bib1.bibx26 bib1.bibx50" id="paren.97"/> and the Pliocene <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx28 bib1.bibx50" id="paren.98"/>, the mid-Miocene can be used to quantify ECS, owing to its relative proximity to modern climate. In this study, we estimated that the ECS of CESM1.2 at the mid-Miocene is 2.9 °C (2.5–3.3 °C, 95 % prediction interval), which is close to the modern-based estimate of ECS of CESM1.2 (2.8 °C, 2.2–3.4 °C, 95 % prediction interval). From this short glance on ECS in CESM1.2, as well as previous work on Miocene ECS <xref ref-type="bibr" rid="bib1.bibx65 bib1.bibx52 bib1.bibx5 bib1.bibx6" id="paren.99"/>, we believe there is motivation in supporting ECS estimates using Miocene evidence, particularly as Miocene modelling is moving towards an official PMIP design. Despite uncertainties in the proxy record or model disagreements, the Pliocene, the warm epoch following the Miocene, has shown that recent warm paleoclimates can provide robust constraint on ECS <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx28 bib1.bibx51" id="paren.100"/>. Additionally, emergent constraint framework can be used to identify disagreements between models, and between model and data <xref ref-type="bibr" rid="bib1.bibx51" id="paren.101"/>. Thus, estimates from the Miocene could either support existing methods using paleoclimate evidence <xref ref-type="bibr" rid="bib1.bibx55 bib1.bibx14" id="paren.102"/>, or be used within a new emergent constraint framework as previously done with emergent constraint frameworks for the LGM <xref ref-type="bibr" rid="bib1.bibx26 bib1.bibx50" id="paren.103"/> or the Pliocene <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx28 bib1.bibx51" id="paren.104"/>.</p>
      <p id="d2e2455">Compared to the proxy record, mid-Miocene simulations are usually either too cold globally, too cold at high latitudes, or both <xref ref-type="bibr" rid="bib1.bibx10" id="paren.105"/>. Over different sensitivity experiments, we find the same issues using CESM1.2, which arise from a weak polar amplification. Simulated high latitudes temperatures underestimate the proxy record by as much as 20 °C, and warm vegetation biomes are more contracted towards low latitudes. However, high CO<sub>2</sub> levels and better representation of high latitude cloud parametrizations produce a better match with high latitude proxy sites and also provided a better match with precipitation proxy reconstruction <xref ref-type="bibr" rid="bib1.bibx1" id="paren.106"/>. Additionally, northern Atlantic sea-surface temperatures reconstructed from coccolith clumped isotopes suggest a more modest warming compared to PI <xref ref-type="bibr" rid="bib1.bibx41" id="paren.107"/>. This would lead to a decrease in the proxy-model bias observed in this region, without overestimating low-latitude ocean temperatures, as inferred from high-CO<sub>2</sub> simulations.</p>
      <p id="d2e2486">A mid-Miocene state close to reality is likely to include a combination of different sensitivities tested in this paper. A continued focus on high-latitude climate feedbacks to enhance polar amplification in models remains important, as this has been a recurring challenge in paleoclimate simulations and could influence projections of future climates. Further research is also needed to assess potential biases and uneven coverage in the proxy record, which can still affect model-proxy comparisons and interpretations of global Miocene climate.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <label>Appendix A</label><title>Ocean model timestep and parametrizations</title>
      <p id="d2e2500">Although changes in ocean parameterization are often required for the numerical stability of paleoclimate simulations, their potential consequences on the climate remain rarely documented. Here, we investigate whether the changes we imposed on our simulations impact our results. These changes are described in Sect. <xref ref-type="sec" rid="Ch1.S2"/> and are namely turning off overflows and tidal mixing, adding a background vertical diffusivity and reducing the timestep of the ocean model.</p>
      <p id="d2e2505">Because these changes are necessary for the stability of our paleoclimate simulations, in particular aspects such as overflows which are hard-coded onto PI geography, we cannot test their impacts onto the mid-Miocene runs. Instead, we apply these configurations to two PI simulations: one, named PI_OcnTimStp, where only the timestep for the computation of the ocean model is reduced from a 63 min timestep to a 41 min timestep; one, named PI_OcnParam, where the full list of changes is applied, including turning off overflows, tidal mixing, adding the background vertical diffusivity coefficient and reducing the ocean timestep. Figure <xref ref-type="fig" rid="FA1"/> summarizes the differences in surface temperatures and AMOC strength relative to the original PI simulation used in our study. We observe colder temperatures by up to 10 °C in localized areas in the North Atlantic ocean where deep-water formation occurs, and a more broad cooling over the Arctic ocean of around 2 °C, as well as a warming of the Southern Ocean by around 1.5 °C in PI_OcnParam relative to PI (Fig. <xref ref-type="fig" rid="FA1"/>A). The temperature differences between PI_OcnTimStp and PI are more negligible (Fig. <xref ref-type="fig" rid="FA1"/>B). The maximum AMOC strength compared to PI (Fig. <xref ref-type="fig" rid="FA1"/>C) weakens by up to 2 Sv in PI_OcnTimStp (Fig. <xref ref-type="fig" rid="FA1"/>D), and by 2.6 Sv in PI_OcnTimStp (Fig. <xref ref-type="fig" rid="FA1"/>E).</p>
      <p id="d2e2521">We highlight that these results are difficult to convey for the mid-Miocene simulations. The changes in ocean parametrizations target specific PI locations which are hard-coded for its geography, but also locations that are connected to stronger circulation. Therefore, we cannot conclude on the impact of the ocean parametrizations on mid-Miocene simulations, because: (1) they have an ocean model initialized uniformly, without any circulation, and (2) they show a PMOC, whereas the colder temperatures found in PI_OcnParam might be a feedback from the weakened AMOC, and the warmer temperatures in the Southern Ocean due to seesaw effect also from the weaker AMOC.</p>
      <p id="d2e2524">One could hypothesize that the ocean parametrizations imposed on CESM1.2 is responsible for the shift to a PMOC in mid-Miocene simulations, if reducing the AMOC strength would push it near a tipping point. However, <xref ref-type="bibr" rid="bib1.bibx19" id="text.108"/> has shown that CESM1.2 is capable of displaying multiple states of equilibrium for its ocean circulation at the Pliocene, including one where both the AMOC and the PMOC co-exists, and we know from Mio_noIS_CAM5 that CESM1.2 can display an ocean with neither an AMOC nor a PMOC. Therefore, it seems unlikely that the complete collapse of the AMOC, and the full-strength PMOC we see in our mid-Miocene simulations is induced by the ocean parametrizations used for the numerical stability.</p><fig id="FA1"><label>Figure A1</label><caption><p id="d2e2533">Differences in surface temperatures between PI and <bold>(A)</bold> PI_OcnTimStp and <bold>(B)</bold> PI_OcnParam, and AMOC strength at <bold>(C)</bold> PI, <bold>(D)</bold> PI_OcnTimStp and <bold>(E)</bold> PI_OcnParam.</p></caption>
        
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f11.png"/>

      </fig>

</app>

<app id="App1.Ch1.S2">
  <label>Appendix B</label><title>Impact of Antarctic ice sheet on heat transport and radiation</title>
      <p id="d2e2567">When implementing an Antarctic ice sheet in our mid-Miocene simulations, we observe a global warming and a particularly pronounced warming over the Arctic region. The climate feedbacks coupling the Antarctic ice sheet to Arctic warming is not straightforward, and beyond the scope of our study. However, we would like to highlight here a few elements to motivate future research on the topic.</p>
      <p id="d2e2570">A first hypothesis was that the ice sheet was somehow causing enhanced heat transport to the Arctic. However, the addition of the ice sheet did not lead to an increase in either oceanic or atmospheric heat transport (Fig. <xref ref-type="fig" rid="FB1"/>). Instead, the difference between atmospheric, oceanic and total heat transport between the simulations is so small, in particular in northern high latitudes, that it suggests local processes are affecting Arctic temperatures.</p>
      <p id="d2e2575">We compare three processes which are related to climate feedbacks in the Arctic region between the simulations in Fig. <xref ref-type="fig" rid="FB2"/>: effective albedo, ice fraction and longwave cloud forcing. When implementing the Antarctic ice sheet in Mio_Ctrl, we observe an overall reduction of surface effective albedo in the Arctic region (over the ocean and land), a reduction in annual Arctic sea-ice and an increase in longwave cloud forcing. All of these elements would favor a warming of the Arctic region compared to Mio_noIS, by either trapping more heat into the system or decreasing the reflection of shortwaves to space.</p>
      <p id="d2e2584">From this brief analysis, we are unable to clearly identify the origin of the warming. Climate feedbacks are known to interact with each other, and in particular in the Arctic, there are notable interactions between cloud and sea-ice feedbacks <xref ref-type="bibr" rid="bib1.bibx35" id="paren.109"/>, as well as a large role from temperature feedbacks <xref ref-type="bibr" rid="bib1.bibx47" id="paren.110"/>, with substantial model differences <xref ref-type="bibr" rid="bib1.bibx4" id="paren.111"/>. Therefore, the problem is left open, and could be relevant for questions of polar amplification in warm paleoclimates.</p><fig id="FB1"><label>Figure B1</label><caption><p id="d2e2599">Atmospheric and oceanic heat transports (PW) per latitude between both Mio_Ctrl and Mio_noIS.</p></caption>
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f12.png"/>

      </fig>

      <fig id="FB2"><label>Figure B2</label><caption><p id="d2e2610">Example of three processes which are tied to climate feedbacks in the Arctic region between Mio_noIS and Mio_Ctrl: effective surface albedo, ice fraction and longwave cloud forcing.</p></caption>
        
        <graphic xlink:href="https://cp.copernicus.org/articles/22/1203/2026/cp-22-1203-2026-f13.png"/>

      </fig>


</app>
  </app-group><notes notes-type="dataavailability"><title>Data availability</title>

      <p id="d2e2627">The temperature and precipitation outputs of the simulations of Table <xref ref-type="table" rid="T2"/> and the vegetation data of Mio_Ctrl are available at <ext-link xlink:href="https://doi.org/10.5281/zenodo.17303799" ext-link-type="DOI">10.5281/zenodo.17303799</ext-link> <xref ref-type="bibr" rid="bib1.bibx49" id="paren.112"/>.</p>
  </notes><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d2e2641">The idea of the study was conceived by MR and AdB. MR performed most simulations, analyses and figures, except the sensitivity simulation on soil color which was performed by TN. The paper was written by MR, AdB, and EB.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d2e2647">The contact author has declared that none of the authors has any competing interests.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d2e2653">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. The authors bear the ultimate responsibility for providing appropriate place names. Views expressed in the text are those of the authors and do not necessarily reflect the views of the publisher.</p>
  </notes><ack><title>Acknowledgements</title><p id="d2e2660">We thank Lauren Burton and one anonymous reviewer for comments that helped improving this paper. We thank Sophia Macarewich and Siva Kattamuri for their advice in preparing the simulations. The computations and data handling were enabled by resources provided by the National Academic Infrastructure for Supercomputing in Sweden (NAISS) partially funded by the Swedish Research Council through grant agreement no. 2022-06725. Getech Group Plc. provided the mid-Miocene paleogeography and support for running the simulations. AdB received funding from the Swedish Research Council grant 2020-04791.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d2e2665">This   research has been supported by the Vetenskapsrådet (grant no. 2020-04791).The publication of this article was funded by the  Swedish Research Council, Forte, Formas, and Vinnova.</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d2e2676">This paper was edited by Heather L. Ford and reviewed by Lauren Burton and one anonymous referee.</p>
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