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<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:oasis="http://docs.oasis-open.org/ns/oasis-exchange/table" xml:lang="en" dtd-version="3.0" article-type="research-article"><?xmltex \makeatother\@nolinetrue\makeatletter?>
  <front>
    <journal-meta><journal-id journal-id-type="publisher">CP</journal-id><journal-title-group>
    <journal-title>Climate of the Past</journal-title>
    <abbrev-journal-title abbrev-type="publisher">CP</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Clim. Past</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1814-9332</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/cp-20-683-2024</article-id><title-group><article-title>Statistical precursor signals for Dansgaard–Oeschger<?xmltex \hack{\break}?> cooling transitions</article-title><alt-title>Statistical precursor signals for DO cooling transitions</alt-title>
      </title-group><?xmltex \runningtitle{Statistical precursor signals for DO cooling transitions}?><?xmltex \runningauthor{T.~Mitsui and N.~Boers}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1 aff2">
          <name><surname>Mitsui</surname><given-names>Takahito</given-names></name>
          <email>takahito321@gmail.com</email>
        <ext-link>https://orcid.org/0000-0002-2825-3996</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff2 aff3">
          <name><surname>Boers</surname><given-names>Niklas</given-names></name>
          
        </contrib>
        <aff id="aff1"><label>1</label><institution>Earth System Modelling, School of Engineering &amp; Design, Technical University of Munich, Munich, Germany</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Potsdam Institute for Climate Impact Research (PIK), Member of the Leibniz Association, <?xmltex \hack{\break}?>P.O. Box 6012 03, 14412 Potsdam, Germany</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Department of Mathematics and Global Systems Institute, University of Exeter, Exeter, UK</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Takahito Mitsui (takahito321@gmail.com)</corresp></author-notes><pub-date><day>22</day><month>March</month><year>2024</year></pub-date>
      
      <volume>20</volume>
      <issue>3</issue>
      <fpage>683</fpage><lpage>699</lpage>
      <history>
        <date date-type="received"><day>10</day><month>June</month><year>2023</year></date>
           <date date-type="rev-request"><day>27</day><month>June</month><year>2023</year></date>
           <date date-type="rev-recd"><day>5</day><month>February</month><year>2024</year></date>
           <date date-type="accepted"><day>8</day><month>February</month><year>2024</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2024 Takahito Mitsui</copyright-statement>
        <copyright-year>2024</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024.html">This article is available from https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024.html</self-uri><self-uri xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024.pdf">The full text article is available as a PDF file from https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d1e106">Given growing concerns about climate tipping points and their risks, it is important to investigate the capability of identifying robust precursor signals for the associated transitions. In general, the variance and short-lag autocorrelations of the fluctuations increase in a stochastically forced system approaching a critical or bifurcation-induced transition, making them theoretically suitable indicators to warn of such transitions. Paleoclimate records provide useful test beds if such a warning of a forthcoming transition could work in practice. The Dansgaard–Oeschger (DO) events are characterized by millennial-scale abrupt climate changes during the glacial period, manifesting most clearly as abrupt temperature shifts in the North Atlantic region. Some previous studies have found such statistical precursor signals for the DO warming transitions. On the other hand, statistical precursor signals for the abrupt DO cooling transitions have not been identified. Analyzing Greenland ice core records, we find robust and statistically significant precursor signals of DO cooling transitions in most of the interstadials longer than roughly 1500 years but not in the shorter interstadials. The origin of the statistical precursor signals is mainly related to so-called rebound events, humps in the temperature observed at the end of interstadial, some decades to centuries prior to the actual transition. We discuss several dynamical mechanisms that give rise to such rebound events and statistical precursor signals.</p>
  </abstract>
    
<funding-group>
<award-group id="gs1">
<funding-source>European Commission</funding-source>
<award-id>TiPES - Tipping Points in the Earth System (820970)</award-id>
<award-id>CriticalEarth - Multiscales and Critical Transitions in the Earth System (956170)</award-id>
</award-group>
<award-group id="gs2">
<funding-source>Bundesministerium für Bildung und Forschung</funding-source>
<award-id>01LS2001A</award-id>
</award-group>
</funding-group>
</article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d1e118">A tipping point is a critical threshold beyond which a system reorganizes, often abruptly and/or irreversibly <xref ref-type="bibr" rid="bib1.bibx38" id="paren.1"/>. Once a tipping point is passed, a system can abruptly transition to an alternative stable or oscillatory state <xref ref-type="bibr" rid="bib1.bibx11" id="paren.2"/>. Empirical and modeling evidence suggests that some components of the Earth system might indeed exhibit tipping behavior, which poses arguably one of the greatest potential risks in the context of ongoing anthropogenic global warming <xref ref-type="bibr" rid="bib1.bibx3 bib1.bibx11" id="paren.3"/>. Paleoclimate evidence supports the fact that abrupt climate changes due to crossing tipping points actually occurred in the past <xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx15 bib1.bibx11" id="paren.4"/>. The Dansgaard–Oeschger events  <xref ref-type="bibr" rid="bib1.bibx27" id="paren.5"/> are one of such past abrupt climate changes during the last glacial period and the focus of this study.</p>
      <p id="d1e136">Tipping point behavior is mathematically classified into three different types <xref ref-type="bibr" rid="bib1.bibx4" id="paren.6"/>. (1) Bifurcation-induced tipping is an abrupt or qualitative change of a system owing to a bifurcation of a stable state (more generally a quasi-static attractor). (2) Noise-induced tipping is an escape from a neighborhood of a quasi-static attractor caused by the action of noisy fluctuations <xref ref-type="bibr" rid="bib1.bibx30" id="paren.7"/>. (3) Rate-induced tipping occurs when a system fails to track a continuously changing quasi-static attractor <xref ref-type="bibr" rid="bib1.bibx4 bib1.bibx87 bib1.bibx63" id="paren.8"/>. In real-world systems, tipping behaviors often result from a combination of several of the above <xref ref-type="bibr" rid="bib1.bibx4" id="paren.9"/>.</p>
      <?pagebreak page684?><p id="d1e151"><?xmltex \hack{\newpage}?>The theory of critical slowing down (CSD) provides a framework to anticipate critical (or bifurcation-induced) transitions <xref ref-type="bibr" rid="bib1.bibx20 bib1.bibx76 bib1.bibx44 bib1.bibx7 bib1.bibx8 bib1.bibx9 bib1.bibx11 bib1.bibx17" id="paren.10"/>. The framework is based on the fact that the stability of a stable state is gradually lost as the system approaches the bifurcation point. Theoretically, the variance of the fluctuations around the fixed point diverges and the autocorrelation with a sufficiently small lag increases toward 1 at the critical point of a codimension-1 bifurcation <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx76 bib1.bibx17" id="paren.11"/>, where the codimension-1 bifurcations are, in simple terms, the bifurcations that can be typically encountered by the change of a single control parameter <xref ref-type="bibr" rid="bib1.bibx82" id="paren.12"/>. Thus, the changes in CSD indicators such as the increase of the variance as well as the autocorrelation can be seen as <italic>statistical precursor signals</italic> (SPSs) of critical transitions. See <xref ref-type="bibr" rid="bib1.bibx26" id="text.13"/> as well as <xref ref-type="bibr" rid="bib1.bibx11" id="text.14"/> for various methods and CSD indicators for anticipating critical transitions.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1" specific-use="star"><?xmltex \currentcnt{1}?><?xmltex \def\figurename{Figure}?><label>Figure 1</label><caption><p id="d1e177">Greenland records from the NGRIP ice core: <bold>(a)</bold> <inline-formula><mml:math id="M1" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and <bold>(b)</bold> <inline-formula><mml:math id="M2" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M3" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] <xref ref-type="bibr" rid="bib1.bibx77 bib1.bibx67" id="paren.15"/>. The interstadial parts longer than <inline-formula><mml:math id="M4" display="inline"><mml:mn mathvariant="normal">1000</mml:mn></mml:math></inline-formula> years are highlighted with gray shades; their numbering is given at the top of each record <xref ref-type="bibr" rid="bib1.bibx67" id="paren.16"/>. The rebound events are indicated by arrows (see Sect. 2.1 for their list). Both records are presented at 20-year resolution. The <inline-formula><mml:math id="M5" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M6" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] record is available only up to DO-24.1. The compositions of GI-23.1 and GI-22, as well as of GI-14 and GI-13, are considered individual long interstadials <xref ref-type="bibr" rid="bib1.bibx19 bib1.bibx67" id="paren.17"/>. The vertical axis for <inline-formula><mml:math id="M7" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M8" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] in <bold>(b)</bold> is reversed to ease visual comparison with the <inline-formula><mml:math id="M9" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O record.</p></caption>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f01.png"/>

      </fig>

      <p id="d1e304">Dansgaard–Oeschger (DO) events are millennial-scale abrupt climate transitions during glacial intervals <xref ref-type="bibr" rid="bib1.bibx27" id="paren.18"/>. They are most clearly imprinted in the <inline-formula><mml:math id="M10" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and calcium ion concentration [Ca<inline-formula><mml:math id="M11" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] records from the Greenland ice cores (Fig. 1) <xref ref-type="bibr" rid="bib1.bibx67 bib1.bibx77" id="paren.19"/>. The <inline-formula><mml:math id="M12" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and [Ca<inline-formula><mml:math id="M13" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] are interpreted as proxies for site temperature and atmospheric circulation changes, respectively. DO warmings occur typically within a few decades and are followed by gradual cooling during relatively warm glacial states termed “interstadials”, before a rapid return to cold states referred to as “stadials”. The amplitude of the abrupt warming transitions ranges from 5 to 16.5 °C (<xref ref-type="bibr" rid="bib1.bibx40" id="altparen.20"/>, and references therein). The Greenland temperatures change concurrently with the North Atlantic temperatures <xref ref-type="bibr" rid="bib1.bibx12 bib1.bibx55" id="paren.21"/>, atmospheric circulation patterns <xref ref-type="bibr" rid="bib1.bibx88" id="paren.22"/>, seawater salinity <xref ref-type="bibr" rid="bib1.bibx32" id="paren.23"/>, sea-ice cover <xref ref-type="bibr" rid="bib1.bibx74" id="paren.24"/>, and the Atlantic Meridional Overturning Circulation (AMOC), as inferred from indices such as <inline-formula><mml:math id="M14" display="inline"><mml:mrow class="chem"><mml:mi mathvariant="normal">Pa</mml:mi><mml:mo>/</mml:mo><mml:mi mathvariant="normal">Th</mml:mi></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M15" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">13</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>C <xref ref-type="bibr" rid="bib1.bibx37" id="paren.25"/>. The combined proxy evidence suggests that the DO events arise from interactions among these components <xref ref-type="bibr" rid="bib1.bibx58 bib1.bibx10" id="paren.26"/>. The prevailing view is that the mode switching of the AMOC plays a principal role in generating DO events <xref ref-type="bibr" rid="bib1.bibx14 bib1.bibx65" id="paren.27"/>, but it remains debated whether the inferred AMOC changes are a driver of DO events or a response to the changes in the atmosphere–ocean–sea-ice system in the North Atlantic, Nordic Seas, and the Arctic <xref ref-type="bibr" rid="bib1.bibx48 bib1.bibx32" id="paren.28"/>. Recently, an increasing number of comprehensive climate models succeeded in simulating DO-like self-sustained oscillations, suggesting that DO events can arise spontaneously from complex interactions between the AMOC, ocean stratification/convection, atmosphere, and sea ice <xref ref-type="bibr" rid="bib1.bibx64 bib1.bibx86 bib1.bibx16 bib1.bibx41 bib1.bibx89 bib1.bibx45 bib1.bibx54" id="paren.29"/>.</p>
      <p id="d1e415">The DO events are considered the archetype of climate tipping behavior <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx15" id="paren.30"/>. Early works found an SPS based on autocorrelation for one specific DO warming – the onset of Bølling–Allerød interstadial <xref ref-type="bibr" rid="bib1.bibx25" id="paren.31"/>. In subsequent works, the existence of SPS for DO warmings was questioned considering that DO warmings are noise induced rather than bifurcation induced <xref ref-type="bibr" rid="bib1.bibx30 bib1.bibx47" id="paren.32"/>. However, a couple of later studies detected SPS for several DO warmings either by ensemble averaging of CSD indicators for many events <xref ref-type="bibr" rid="bib1.bibx22" id="paren.33"/> or by using wavelet-transform techniques focusing on a specific frequency band <xref ref-type="bibr" rid="bib1.bibx73 bib1.bibx7" id="paren.34"/>. On the other hand, it has so far not been shown whether DO coolings are preceded by characteristic CSD-based precursor signals as well.</p>
      <p id="d1e433">Recent studies have inferred that the AMOC is currently at its weakest in at least a millennium <xref ref-type="bibr" rid="bib1.bibx66 bib1.bibx18" id="paren.35"/> (see also <xref ref-type="bibr" rid="bib1.bibx39" id="text.36"/> for possible uncertainties). The declining AMOC trend is projected to continue in the coming century, although the projections of the AMOC strength in the next 100 years are model dependent <xref ref-type="bibr" rid="bib1.bibx56" id="paren.37"/>. Furthermore, the studies applying CSD indicators to observed AMOC fingerprints <xref ref-type="bibr" rid="bib1.bibx8 bib1.bibx5 bib1.bibx29" id="paren.38"/> as well as a long-term reconstruction of the Atlantic multidecadal variability <xref ref-type="bibr" rid="bib1.bibx59" id="paren.39"/> suggest that the AMOC stability may have declined and the AMOC might be approaching a dangerous tipping point. In this context, it is important to investigate whether CSD-based precursor signals can be detected for the DO cooling transitions as well, supposing that the DO events reflect past AMOC changes. However, of course, predictability of past events does not necessarily imply any predictability in the future, especially given that the recent AMOC weakening is presumably driven by global warming and is thus from a mechanistic point of view different from past AMOC weakenings in the glacial period.</p>
      <p id="d1e451">In this study we report SPS for DO cooling transitions recorded in <inline-formula><mml:math id="M16" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and <inline-formula><mml:math id="M17" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M18" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] <xref ref-type="bibr" rid="bib1.bibx77 bib1.bibx67" id="paren.40"/> from three Greenland ice cores: the North Greenland Ice Core Project (NGRIP), the Greenland Ice Core Project (GRIP), and the Greenland Ice Sheet Project 2 (GISP2) (see Fig. 1 for NGRIP). The important source of observed SPS stems from so-called rebound events, humps in the temperature proxy occurring at the end of interstadials, some decades to centuries prior to the transition <xref ref-type="bibr" rid="bib1.bibx19" id="paren.41"/>. When CSD indicators such as variance or lag-1 autocorrelation are used for anticipating a tipping point, we conventionally assume that a system gradually approaches a bifurcation point. However, if DO cycles are spontaneous oscillations as suggested in the comprehensive climate models (see above), in a strict sense there might not be any bifurcation occurring around the timings of the abrupt<?pagebreak page685?> transition in the DO cycles. Nevertheless, with conceptual models of DO cycles, we demonstrate that CSD indicators may show precursor signals for abrupt transitions due to particular dynamics near a critical point or a critical manifold.</p>
      <p id="d1e494">The remainder of this paper is organized as follows. In Sect. 2, the data and method are described. In Sect. 3, we identify robust and statistically significant SPSs for several DO cooling transitions following interstadials with sufficient data length and show that rebound events prior to cooling transition are the source of observed SPSs. In Sect. 4 we discuss the results by using conceptual models. A summary is given in Sect. 5.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Data and methods</title>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Greenland ice core records</title>
      <p id="d1e512">We explore CSD-based precursor signals for DO cooling transitions recorded in <inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and <inline-formula><mml:math id="M20" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M21" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] <xref ref-type="bibr" rid="bib1.bibx77 bib1.bibx67" id="paren.42"/> from three Greenland ice cores: NGRIP, GRIP, and GISP2 (see Fig. 1 for NGRIP). Multiple records are used for a robust assessment because each has regional fluctuations as well as proxy- and ice-core-dependent uncertainties. The six records have been synchronized and are given at 20-year resolution <xref ref-type="bibr" rid="bib1.bibx77 bib1.bibx67" id="paren.43"/>. They continuously span the last 104 kyr b2k (kiloyears before 2000 CE), beyond which only NGRIP <inline-formula><mml:math id="M22" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O is available up to a part of the Eemian Interglacial. In addition, we use a version of the NGRIP <inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and dust records at 5 cm depth resolution <xref ref-type="bibr" rid="bib1.bibx33 bib1.bibx35 bib1.bibx72" id="paren.44"/> in order to check the dependence of results on temporal resolutions, with the caveat that these high-resolution records span only the last 60 kyr.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2" specific-use="star"><?xmltex \currentcnt{2}?><?xmltex \def\figurename{Figure}?><label>Figure 2</label><caption><p id="d1e583">Analysis of CSD-based precursor signals of abrupt DO cooling transitions, for the first six interstadials of NGRIP <inline-formula><mml:math id="M24" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O, from 115 ka to 74 kyr b2k. Top row: interstadials longer than <inline-formula><mml:math id="M25" display="inline"><mml:mn mathvariant="normal">1000</mml:mn></mml:math></inline-formula> years (blue). The cooling transition and stadial parts are shown in gray <xref ref-type="bibr" rid="bib1.bibx67" id="paren.45"/>. Nonlinear trends are calculated with the locally weighted scatterplot smoothing (LOESS) (red). The smoothing span <inline-formula><mml:math id="M26" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> that defines the fraction of data points involved in the local regression is set to <inline-formula><mml:math id="M27" display="inline"><mml:mn mathvariant="normal">50</mml:mn></mml:math></inline-formula> % of each interstadial length. The rebound events are indicated by arrows (see Sect. 2.1). Second row: residuals (green) resulting from subtracting the nonlinear trends (red) from the records (blue). Third row: variance estimate in rolling windows (black) with size equal to 50 % of each interstadial length. Values are plotted at the right edge of each rolling window. The linear trend is shown by a solid red line for <inline-formula><mml:math id="M28" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>, by a dashed red line for <inline-formula><mml:math id="M29" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.05</mml:mn><mml:mo>&lt;</mml:mo><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>, and by a dotted line for <inline-formula><mml:math id="M30" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>. Fourth row: same as third row but for the lag-1 autocorrelation (i.e., a lag of 20 years).</p></caption>
          <?xmltex \igopts{width=483.69685pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f02.png"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3" specific-use="star"><?xmltex \currentcnt{3}?><?xmltex \def\figurename{Figure}?><label>Figure 3</label><caption><p id="d1e670">Same as Fig. 2 but for the next six interstadials, from 74 ka to 12 kyr b2k.</p></caption>
          <?xmltex \igopts{width=483.69685pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f03.png"/>

        </fig>

      <p id="d1e680">We follow the classification of interstadials and stadials and associated timings of DO warming and cooling transitions by <xref ref-type="bibr" rid="bib1.bibx67" id="text.46"/>, where Greenland interstadials (stadials) are labeled with “GI” (“GS”) with a few exceptions below. A <italic>rebound event</italic> is an abrupt warming often observed before an interstadial abruptly ends <xref ref-type="bibr" rid="bib1.bibx19" id="paren.47"/> (arrows in Figs. 1, 2, and 3). Generally, a long interstadial accompanies a long rebound event (their durations are correlated with <inline-formula><mml:math id="M31" display="inline"><mml:mrow><mml:msup><mml:mi>R</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.95</mml:mn></mml:mrow></mml:math></inline-formula>, <xref ref-type="bibr" rid="bib1.bibx19" id="altparen.48"/>). In <xref ref-type="bibr" rid="bib1.bibx19" id="text.49"/> and <xref ref-type="bibr" rid="bib1.bibx67" id="text.50"/>, GI-14 and the subsequent interstadial, GI-13, are seen as one long interstadial, with GI-13 considered to be a strongly expressed rebound event ending GI-14 because the changes in <inline-formula><mml:math id="M32" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and <inline-formula><mml:math id="M33" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M34" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] during the quasi-stadial GS-14 do not reach the baseline levels of adjacent stadials. Similarly GI-23.1 and GI-22 are also seen as one long interstadial, and GI-22 is<?pagebreak page686?> regarded as a rebound event (and GS-23.1 as quasi-stadial) <xref ref-type="bibr" rid="bib1.bibx19 bib1.bibx67" id="paren.51"/>. GI-20a is also recognized as a rebound event in <xref ref-type="bibr" rid="bib1.bibx67" id="text.52"/>. Given that the rebound events are warmings following a colder spell during interstadial conditions that does not reach the stadial levels <xref ref-type="bibr" rid="bib1.bibx67" id="paren.53"/>, we regard the following nine epochs as rebound-type events: GI-8a, the hump at the end of GI-11 (42 240–<inline-formula><mml:math id="M35" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">42</mml:mn><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mn mathvariant="normal">500</mml:mn></mml:mrow></mml:math></inline-formula> yr b2k), GI-12a, GI-13, the hump at the end of GI-16.1 (56 500–<inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">56</mml:mn><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mn mathvariant="normal">900</mml:mn></mml:mrow></mml:math></inline-formula> yr b2k), GI-20a, GI-21.1c-b-a (two warming transitions), GI-22, and GI-25a. When we examine the effect of rebound events on our results, we exclude the entire parts including the cold spells prior to the rebound events.</p>
      <p id="d1e787">The start (warming) and end (cooling) of each DO event are identified in 20-year resolution based on both <inline-formula><mml:math id="M37" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O and [Ca<inline-formula><mml:math id="M38" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] in <xref ref-type="bibr" rid="bib1.bibx67" id="text.54"/>. The estimated uncertainty of event timing varies from event to event. We remove the 2<inline-formula><mml:math id="M39" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula> uncertainty range of the event timing (40 to 400 years) estimated in <xref ref-type="bibr" rid="bib1.bibx67" id="text.55"/> from our calculation of CSD indicators. It effectively excludes parts of the transitions themselves from the calculation of the CSD indicators. Since the calculation of CSD indicators requires a minimum length of data points, we mainly focus on interstadials longer than 1000 years after removing 2<inline-formula><mml:math id="M40" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula> uncertainty ranges of the transition timings, using 20-year resolution data (Table S1 in the Supplement). This results in 12 DO interstadials to be<?pagebreak page687?> investigated (Fig. 1, gray shades). We deal with the interstadials shorter than 1000 years but longer than 300 years using high-resolution records in Sect. 3.2.</p>
</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>Statistical indicators of critical slowing down</title>
      <p id="d1e842">Based on the theory of critical slowing down (CSD), we posit that the stability of a dynamical system perturbed by noise is gradually lost as the system approaches a bifurcation point <xref ref-type="bibr" rid="bib1.bibx11 bib1.bibx76 bib1.bibx17" id="paren.56"/>. For the fold bifurcation (also known as the saddle-node bifurcation), the variance of the fluctuations around a local stable state diverges and the autocorrelation function of the fluctuations increases toward <inline-formula><mml:math id="M41" display="inline"><mml:mn mathvariant="normal">1</mml:mn></mml:math></inline-formula> at any lag <inline-formula><mml:math id="M42" display="inline"><mml:mi mathvariant="italic">τ</mml:mi></mml:math></inline-formula>. The same is true for the transcritical as well as the pitchfork bifurcation <xref ref-type="bibr" rid="bib1.bibx17" id="paren.57"/>. For the Hopf bifurcation, the variance increases, but the autocorrelation function of the form <inline-formula><mml:math id="M43" display="inline"><mml:mrow><mml:mi>C</mml:mi><mml:mo>(</mml:mo><mml:mi mathvariant="italic">τ</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi mathvariant="italic">ν</mml:mi><mml:mo>|</mml:mo><mml:mi mathvariant="italic">τ</mml:mi><mml:mo>|</mml:mo></mml:mrow></mml:msup><mml:mi>cos⁡</mml:mi><mml:mi mathvariant="italic">ω</mml:mi><mml:mi mathvariant="italic">τ</mml:mi></mml:mrow></mml:math></inline-formula> may increase or decrease depending on <inline-formula><mml:math id="M44" display="inline"><mml:mi mathvariant="italic">τ</mml:mi></mml:math></inline-formula>, where <inline-formula><mml:math id="M45" display="inline"><mml:mrow><mml:mi mathvariant="italic">ν</mml:mi><mml:mo>(</mml:mo><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M46" display="inline"><mml:mrow><mml:mo>±</mml:mo><mml:mi mathvariant="italic">ω</mml:mi><mml:mi>i</mml:mi></mml:mrow></mml:math></inline-formula> are the real and imaginary parts of the complex eigenvalues of the Jacobian matrix of the local linearized system <xref ref-type="bibr" rid="bib1.bibx17" id="paren.58"/>. Nevertheless, the autocorrelation function <inline-formula><mml:math id="M47" display="inline"><mml:mrow><mml:mi>C</mml:mi><mml:mo>(</mml:mo><mml:mi mathvariant="italic">τ</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> increases for sufficiently small <inline-formula><mml:math id="M48" display="inline"><mml:mi mathvariant="italic">τ</mml:mi></mml:math></inline-formula>. For discrete time series, we follow previous studies and calculate the lag-1 autocorrelation corresponding to a minimal <inline-formula><mml:math id="M49" display="inline"><mml:mi mathvariant="italic">τ</mml:mi></mml:math></inline-formula>. These characteristics can be used to anticipate abrupt transitions caused by codimension-1 bifurcations. Promisingly previous studies show that these CSD indicators and related measures can indeed anticipate simulated AMOC collapses <xref ref-type="bibr" rid="bib1.bibx13 bib1.bibx42 bib1.bibx49 bib1.bibx36" id="paren.59"/>.</p>
      <p id="d1e970">Prior to calculating CSD indicators, we estimate the local stable state by using a local regression method called the <italic>locally weighted scatterplot smoothing</italic> (LOESS) <xref ref-type="bibr" rid="bib1.bibx24 bib1.bibx26" id="paren.60"/>. In this approach, the time series is seen as a scatter plot and fitted locally by a polynomial function, giving more weight to points near the point whose response is being estimated and less weight to points further away. Here the polynomial degree is set to 1; i.e., the smoothing is performed with the local linear fit. Nevertheless, the LOESS provides nonlinear smoothed curves. The smoothing span parameter <inline-formula><mml:math id="M50" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> that defines the fraction of data points<?pagebreak page688?> involved in the local regression is set to 50 % of each interstadial length in a demonstration case, but we examine the dependence of results on <inline-formula><mml:math id="M51" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> over the range 20 %–70 %. The difference between the record and the smoothed one gives the residual fluctuations. The CSD indicators, i.e., variance and lag-1 autocorrelation, are calculated for the residuals over a rolling window. The size <inline-formula><mml:math id="M52" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> of this rolling window is set to 50 % of each interstadial length in the demonstration case. To test the robustness, this is changed over the range 20 %–60 %.</p>
      <p id="d1e1000">The statistical significance of precursor signals of critical transitions, in terms of positive trends of CSD indicators, is assessed by hypothesis testing <xref ref-type="bibr" rid="bib1.bibx80 bib1.bibx26 bib1.bibx73 bib1.bibx7" id="paren.61"/>. We consider as null model a stationary stochastic process with preserved variance and autocorrelation. The <inline-formula><mml:math id="M53" display="inline"><mml:mi>n</mml:mi></mml:math></inline-formula> surrogate data are prepared from the original residual series by the phase-randomization method, thus preserving the variance and autocorrelation function of the original time series via the Wiener–Khinchin theorem. Here we take <inline-formula><mml:math id="M54" display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1000</mml:mn></mml:mrow></mml:math></inline-formula>. The linear trend (<inline-formula><mml:math id="M55" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">o</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) of the CSD indicator for the original time series and the linear trends (<inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) of CSD indicators for the surrogate data are calculated. We consider the precursor signal of the original time series statistically significant at the 5 % level if the probability of <inline-formula><mml:math id="M57" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub><mml:mo>&gt;</mml:mo><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">o</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M58" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula> value) is less than 0.05. The significance level of 0.05 is conservative given that some works analyzing ecological or paleo-data adopt the significance level of 0.1 <xref ref-type="bibr" rid="bib1.bibx26 bib1.bibx81" id="paren.62"/>.</p>
</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Results</title>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Characteristic precursor signals of DO coolings</title>
      <p id="d1e1092">As CSD indicators we consider the variance and lag-1 autocorrelation, calculated in rolling windows across each interstadial. The 12 interstadials longer than 1000 years are magnified in Figs. 2 and 3 (top rows, blue) for the NGRIP <inline-formula><mml:math id="M59" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O record. See Figs. S1–S10 in the Supplement for the other records. For each interstadial, the nonlinear trend is estimated using LOESS smoothing (Figs. 2 and 3, top row, red). In this case the smoothing span <inline-formula><mml:math id="M60" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> that defines the fraction of data points involved in the local regression is set to <inline-formula><mml:math id="M61" display="inline"><mml:mn mathvariant="normal">50</mml:mn></mml:math></inline-formula> % of each interstadial length. Gaussian kernel smoothing gives similar results. The difference between the record and the nonlinear trend gives the approximately stationary residual fluctuations (second row). The CSD indicators are calculated from the residual series over a rolling window. In Figs. 2 and 3 the rolling window size <inline-formula><mml:math id="M62" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> is set to 50 % of each interstadial length (a default value in <xref ref-type="bibr" rid="bib1.bibx25" id="altparen.63"/>). The smoothing span <inline-formula><mml:math id="M63" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> and the rolling window size <inline-formula><mml:math id="M64" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> are taken as fractions of individual interstadial length because timescales of local fluctuations (such as the duration of rebound events) change with the entire duration of interstadial. We examine the dependence of the results on <inline-formula><mml:math id="M65" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M66" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> as part of our robustness tests.</p>
      <p id="d1e1159">The variance is plotted in the third row of Figs. 2 and 3. Positive trends in the variance are observed for 9 out of 12 interstadials; the individual trends are statistically significant in 6 out of 12 cases (<inline-formula><mml:math id="M67" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>), based on a null model assuming the same overall variance and autocorrelation, constructed by producing surrogates with randomized Fourier phases. The lag-1 autocorrelation is also plotted for the same data in the bottom row. Positive trends in lag-1 autocorrelation are observed for 10 out of 12 interstadials but are statistically significant only in two cases (<inline-formula><mml:math id="M68" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>). Just a positive trend without significance cannot be considered a reliable SPS, but if one indicator has a significantly positive trend, the other indicator with a consistently positive trend may at least support the conclusion (e.g., GI-19.2 and GI-14-13 in Fig. 3). In several cases (GI-24.2, GI-21.1, GI-16.1, GI-14-13, and GI-12), the lag-1 autocorrelation first decreases and then increases. The initial decreases, harming monotonic increases of CSD indicators, might reflect a memory of the preceding DO warming transition. On the other hand, the drastic increases in both indicators near the end of the interstadials reflect the rebound events (arrows in Figs. 2 and 3). We obtain similar results for the other ice core records  (Figs. S1–S10). While we observe a number of positive trends for all the records, the number of statistically significant trends detected depends on the record and CSD indicator (Table S2).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4" specific-use="star"><?xmltex \currentcnt{4}?><?xmltex \def\figurename{Figure}?><label>Figure 4</label><caption><p id="d1e1188">Detection of precursor signals of DO cooling transitions for different interglacials, different proxy variables, different ice cores, and different CSD indicators. <bold>(a, b)</bold> Robustness analysis of precursor signals with respect to the smoothing span and the rolling window size (% of interstadial length): the case of the GI-25 interstadial from the NGRIP <inline-formula><mml:math id="M69" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O record. The CSD indicator is the variance in <bold>(a)</bold> and the lag-1 autocorrelation in <bold>(b)</bold>. Cross marks (x) indicate statistically significant positive trends of the respective CSD indicator (<inline-formula><mml:math id="M70" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>) based on a phase surrogate test (see Sect. 2.2), small open circles (o) indicate barely significant positive trends (<inline-formula><mml:math id="M71" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.05</mml:mn><mml:mo>&lt;</mml:mo><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>), and cells are left blank if <inline-formula><mml:math id="M72" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>. <bold>(c)</bold> Durations of interstadials longer than 1000 years (Table S1). The dashed line indicates 1500 years <bold>(d)</bold>. Robustness of finding precursor signals for DO cooling transitions. The color indicates the number of significant (<inline-formula><mml:math id="M73" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>) positive trends in each of the 30 sets of the smoothing spans and the rolling window sizes as in <bold>(a)</bold> and <bold>(b)</bold>. For the cases of gray-shaded cells, the data are not publicly available.</p></caption>
          <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f04.png"/>

        </fig>

      <p id="d1e1284">We check the robustness of our results against changing smoothing span <inline-formula><mml:math id="M74" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> and rolling window size <inline-formula><mml:math id="M75" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx26" id="paren.64"/>. We calculate the <inline-formula><mml:math id="M76" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula> value for the trend of each indicator changing the smoothing span between 20 % and 70 % of interstadial length (in steps of <inline-formula><mml:math id="M77" display="inline"><mml:mn mathvariant="normal">10</mml:mn></mml:math></inline-formula> %) and the rolling window size between 20 % and 60 % (also in <inline-formula><mml:math id="M78" display="inline"><mml:mn mathvariant="normal">10</mml:mn></mml:math></inline-formula> % steps). This yields a <inline-formula><mml:math id="M79" display="inline"><mml:mrow><mml:mn mathvariant="normal">6</mml:mn><mml:mo>×</mml:mo><mml:mn mathvariant="normal">5</mml:mn></mml:mrow></mml:math></inline-formula> matrix for the <inline-formula><mml:math id="M80" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula> values. Example results for GI-25 and <inline-formula><mml:math id="M81" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O are shown in Fig. 4a (variance) and 4b (lag-1 autocorrelation). The cross mark (x) indicates significant positive trends (<inline-formula><mml:math id="M82" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>) and the small open circle (o) indicates positive trends that are significant at the 10 % level but not at the 5 % level. Full results for the 12 interstadials, six records, and two CSD indicators are shown in   Figs. S11–S22. We consider positive trends in CSD indicators, i.e., the SPS of the transition, to be overall robust if we obtain significant positive trends (<inline-formula><mml:math id="M83" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>) for more than half (<inline-formula><mml:math id="M84" display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">15</mml:mn></mml:mrow></mml:math></inline-formula>) of the 30 parameter sets.</p>
      <p id="d1e1391">The robustness analysis is performed for all the long interstadials of the six records and the two CSD indicators (Fig. 4d). Among the 12 interstadials, we find at least one robust SPS for eight interstadials (GI-25, GI-23.1, GI-21.1, GI-20, GI-19.2, GI-14, GI-12, and GI-8) and multiple robust SPSs for six (GI-25, GI-23.1, GI-21.1, GI-14, GI-12, and GI-8). If the data series is a stationary stochastic process, the probability of spuriously observing a robust SPS is estimated to be <inline-formula><mml:math id="M85" display="inline"><mml:mn mathvariant="normal">5</mml:mn></mml:math></inline-formula> % (Appendix A). In this case, the probability of detecting more than two robust SPSs from 12 independent stationary samples (i.e., from each row in Fig. 4d) by chance is only <inline-formula><mml:math id="M86" display="inline"><mml:mrow><mml:mo>∼</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula> %. Thus, the risk of obtaining the<?pagebreak page689?> results by chance is quite low. For each interstadial, the detection of robust SPSs depends on the proxy and core. This is possibly because different proxies from different cores are contaminated by different types and magnitudes of noise (e.g., <inline-formula><mml:math id="M87" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O may record local fluctuations of temperatures and <inline-formula><mml:math id="M88" display="inline"><mml:mrow><mml:msub><mml:mi>log⁡</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>[Ca<inline-formula><mml:math id="M89" display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>+</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula>] turbulent fluctuations of local wind circulations). Robust SPSs are observed for most interstadials longer than roughly 1500 years (GI-25, GI-23.1, GI-21.1, GI-20, GI-19.2, GI-14, GI-12, and GI-8 except GI-1) but not for the other interstadials, shorter than roughly 1500 years (compare Figs. 4c and 4d).</p>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Further sensitivity analyses</title>
      <p id="d1e1453">We examine how much the rebound events affect the detection of CSD-based SPS. For this purpose CSD indicators are again calculated excluding the rebound events and their preceding cold spells (see Sect. 2.1). While eight interstadials (GI-25, GI-23.1, GI-21.1, GI-20, GI-16, GI-14, GI-12, and GI-8) exhibit robust SPS with the rebound events included,<?pagebreak page690?> only four interstadials (GI-23.1, GI-14, GI-12, and GI-8) exhibit robust SPS without the rebound events (Fig. S23). The rebound events should hence be considered important, sometimes indispensable, sources for SPS of DO coolings.</p>
      <p id="d1e1456">We also examine the dependence of the results on the time resolution of the data. Here we use a high-resolution (5 cm depth) <inline-formula><mml:math id="M90" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O record <xref ref-type="bibr" rid="bib1.bibx33 bib1.bibx35" id="paren.65"/> and a dust record <xref ref-type="bibr" rid="bib1.bibx72" id="paren.66"/> from the NGRIP over the last 60 kyr. Since the data in these records are non-uniform in time, they are linearly resampled every 5 years before calculating CSD indicators. We focus on 11 interstadials longer than 300 years in order to have enough data points. For the dust record, three interstadials (GI-15, GI-8, and GI-7) are excluded from the analysis because the original data have long parts of missing values. The CSD indicators, calculated with a smoothing span of <inline-formula><mml:math id="M91" display="inline"><mml:mrow><mml:mi mathvariant="italic">α</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">50</mml:mn></mml:mrow></mml:math></inline-formula> % and rolling windows of <inline-formula><mml:math id="M92" display="inline"><mml:mrow><mml:mi>W</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">50</mml:mn></mml:mrow></mml:math></inline-formula> %, are shown in Figs. S24–S27. Through the robustness analyses with respect to <inline-formula><mml:math id="M93" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M94" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula>, we find at least one robust SPS for three out of 11 interstadials (Fig. S28). The robust SPSs for GI-14-13 and GI-12 from the high-resolution records are consistent with those from the 20-year resolution records. Moreover for GI-1, the high-resolution <inline-formula><mml:math id="M95" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O record exhibits a robust SPS in terms of lag-1 autocorrelation, although the 20-year resolution record does not. Robust SPSs have not been observed again for interstadials shorter than roughly 1500 years (Figs. S28 and 4).</p>

      <?xmltex \floatpos{p}?><fig id="Ch1.F5" specific-use="star"><?xmltex \currentcnt{5}?><?xmltex \def\figurename{Figure}?><label>Figure 5</label><caption><p id="d1e1528">Four potential dynamical mechanisms for the DO cooling transitions. <bold>(a)</bold> Fold bifurcation mechanism. The time series <inline-formula><mml:math id="M96" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> for decreasing <inline-formula><mml:math id="M97" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> (blue). The green lines show the stable (solid) and unstable (dashed) fixed points. <bold>(b, c)</bold> Stochastic slow–fast oscillation mechanism of a FitzHugh–Nagumo-type model. An example time series <inline-formula><mml:math id="M98" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is shown in <bold>(b)</bold> and the phase space trajectory (blue) in <bold>(c)</bold>; the <inline-formula><mml:math id="M99" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> nullcline, i.e., the critical manifold, is shown in green. The <inline-formula><mml:math id="M100" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> nullcline is shown in dashed magenta. <bold>(d, e)</bold> Hopf bifurcation mechanism. An example time series <inline-formula><mml:math id="M101" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is shown in <bold>(d)</bold> as a function of <inline-formula><mml:math id="M102" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. Stable (black, solid) and unstable (magenta, dashed) fixed points are also shown. The corresponding phase space trajectory (<inline-formula><mml:math id="M103" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>) for decreasing <inline-formula><mml:math id="M104" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> is shown in <bold>(e)</bold> in blue. The critical manifold is shown in green. <bold>(f, g)</bold> Mixed-mode oscillation mechanism. An example time series <inline-formula><mml:math id="M105" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is shown in <bold>(f)</bold> and the corresponding phase space trajectory in <bold>(g)</bold>. The magenta dot is the saddle point with a stable manifold in the direction of the black segment; the trajectory is spiralling around it.</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f05.png"/>

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      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><?xmltex \currentcnt{6}?><?xmltex \def\figurename{Figure}?><label>Figure 6</label><caption><p id="d1e1702">Rate dependence of CSD indicators for the fold bifurcation in the Stommel model. Its parameter <inline-formula><mml:math id="M106" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is decreased from <inline-formula><mml:math id="M107" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula> to <inline-formula><mml:math id="M108" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.4</mml:mn></mml:mrow></mml:math></inline-formula> across the fold bifurcation point <inline-formula><mml:math id="M109" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.25</mml:mn></mml:mrow></mml:math></inline-formula> with a small rate <inline-formula><mml:math id="M110" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">μ</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">600</mml:mn></mml:mrow></mml:math></inline-formula> (left column) or with a 10 times larger rate <inline-formula><mml:math id="M111" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi mathvariant="italic">μ</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">60</mml:mn></mml:mrow></mml:math></inline-formula> (right column). <bold>(a, b)</bold> Dynamical variable <inline-formula><mml:math id="M112" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> before the tipping (blue) and after the tipping (gray). The time interval of sampling <inline-formula><mml:math id="M113" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> value is 0.1 in both cases. The solid black lines and the dashed lines show the stable equilibria and the unstable equilibrium, respectively. <bold>(c, d)</bold> Residuals (green) resulting from subtracting the nonlinear trends (red) from the records (blue). <bold>(e, f)</bold> Variance estimate in rolling windows (black) with size equal to 50 % of the signal before tipping. Values are plotted at the right edge of each rolling window. The linear trend is shown by a solid red line for <inline-formula><mml:math id="M114" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula> and by a dotted line for <inline-formula><mml:math id="M115" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>. <bold>(g, h)</bold> Same as <bold>(e, f)</bold> but for the lag-1 autocorrelation. Significant increases of CSD indicators are unlikely to be observed when the parameter changes rapidly. Typical cases are shown here. Details of trajectories and the changes of CSD indicators depend on noise realizations.</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://cp.copernicus.org/articles/20/683/2024/cp-20-683-2024-f06.png"/>

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</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Possible dynamical mechanisms</title>
      <p id="d1e1897">We detected robust precursor signals of DO cooling transitions for most interstadials longer than roughly 1500 years but not for shorter interstadials. The results suggest that long interstadials, the existence of rebound events, and the presence of SPS for the DO cooling transitions are all related (except for GI-19.2, which has no noticeable rebound event). These aspects may be related to generic properties of nonlinear dynamical systems. On the basis of conceptual mathematical models, we discuss four possible dynamical mechanisms leading to the precursor signals of DO cooling transitions. In three of four mechanisms, oscillations such as the rebound events can systematically arise prior to the abrupt cooling transitions. These modeling results justify the inclusion of the rebound events in the search for precursor signals presented above. Unless otherwise mentioned, details on model parameters as well as the hysteresis experiments conducted are given in Appendix B. <list list-type="order"><list-item>
      <p id="d1e1902"><italic>The fold bifurcation mechanism</italic>. Since the pioneering work by <xref ref-type="bibr" rid="bib1.bibx78" id="text.67"/>, the AMOC is considered to exhibit bistability depending on the background condition <xref ref-type="bibr" rid="bib1.bibx65" id="paren.68"/>. The bistability of the AMOC strength <inline-formula><mml:math id="M116" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> may be conceptually modeled by a double-fold bifurcation model: <inline-formula><mml:math id="M117" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M118" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> has two fold points such as <inline-formula><mml:math id="M119" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M120" display="inline"><mml:mrow><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. Here we take the quadratic from <inline-formula><mml:math id="M121" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, but the following arguments are qualitatively the same for <inline-formula><mml:math id="M122" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>. The parameter <inline-formula><mml:math id="M123" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> represents the surface salinity flux (i.e., negative freshwater flux), and <inline-formula><mml:math id="M124" display="inline"><mml:mrow><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> denotes white Gaussian noise. The unperturbed model for <inline-formula><mml:math id="M125" display="inline"><mml:mrow><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> has equilibria on an S-shaped curve: <inline-formula><mml:math id="M126" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> (Fig. 5a, green). The state <inline-formula><mml:math id="M127" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> initially on the upper stable branch jumps down to the lower stable branch as <inline-formula><mml:math id="M128" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> decreases across the fold bifurcation point at <inline-formula><mml:math id="M129" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.25</mml:mn></mml:mrow></mml:math></inline-formula>. The variance and the autocorrelation of the local fluctuations (i.e., CSD indicators) increase as <inline-formula><mml:math id="M130" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> slowly approaches the fold bifurcation point since the restoring rate toward the stable state decreases, as shown in Fig. 6a <xref ref-type="bibr" rid="bib1.bibx76 bib1.bibx11" id="paren.69"/>.</p></list-item><list-item>
      <p id="d1e2165"><italic>Stochastic slow–fast oscillation mechanism</italic>. The FitzHugh–Nagumo (FHN) system is a prototypical model for slow–fast oscillations and excitability <xref ref-type="bibr" rid="bib1.bibx34 bib1.bibx62" id="paren.70"/>. It is often invoked for conceptual models of DO oscillations <xref ref-type="bibr" rid="bib1.bibx68 bib1.bibx46 bib1.bibx71 bib1.bibx61 bib1.bibx50 bib1.bibx69 bib1.bibx86" id="paren.71"/>. An FHN-type model of DO oscillations can be obtained by introducing a slow variable <inline-formula><mml:math id="M131" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> into the fold bifurcation model: <inline-formula><mml:math id="M132" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M133" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>y</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M134" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M135" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are timescale parameters (<inline-formula><mml:math id="M136" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>≪</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>). Invoking the salt-oscillator hypothesis for DO oscillations suggested by the comprehensive climate model simulations that are successful in reproducing DO cycles <xref ref-type="bibr" rid="bib1.bibx85" id="paren.72"/>, we may interpret <inline-formula><mml:math id="M137" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> as the surface mixed-layer salinity in the northern North Atlantic and Labrador Sea, which gradually decreases (increases) when the AMOC intensity <inline-formula><mml:math id="M138" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> is strong (weak).</p>
      <?pagebreak page692?><p id="d1e2338">Here we consider the case that the system is excitable. For example, for <inline-formula><mml:math id="M139" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.26</mml:mn></mml:mrow></mml:math></inline-formula>, the unperturbed system has a stable equilibrium near the upper fold point of the S-shaped critical manifold, <inline-formula><mml:math id="M140" display="inline"><mml:mrow><mml:mo mathvariant="italic">{</mml:mo><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo><mml:mo>∈</mml:mo><mml:msup><mml:mi mathvariant="double-struck">R</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mo>|</mml:mo><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo mathvariant="italic">}</mml:mo></mml:mrow></mml:math></inline-formula>  (Fig. 5c, green), but the dynamical noise <inline-formula><mml:math id="M141" display="inline"><mml:mrow><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> enables the escape from the barely stable equilibrium and sustains stochastic oscillations (Fig. 5b and c, blue). Due to the timescale separation (<inline-formula><mml:math id="M142" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>≪</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), the oscillations occur along the attracting parts of the critical manifold (Fig. 5c). Because <inline-formula><mml:math id="M143" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> is much slower than <inline-formula><mml:math id="M144" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>, the dynamics of <inline-formula><mml:math id="M145" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> is similar to the dynamics of the fold bifurcation model with slowly changing <inline-formula><mml:math id="M146" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula>. As a result, SPS can be effectively observed near the upper fold point of the critical manifold  (Fig. S29). However, this example is not rigorous bifurcation-induced tipping. In the example of an excitable system (Fig. 5b and c), the underlying system always has a weakly stable fixed point, and no true bifurcation leading to CSD occurs. In fact, the actual tipping in this case is noise induced. However, we can effectively observe the SPS in CSD indicators in this case as well, since the system would in each cyclic iteration move from more stable to less stable conditions until it finally tips to initiate the next cycle, and this partial decrease in stability is imprinted in the CSD indicators (Fig. S29). The increase of the variance prior to the transitions in the FHN model is also reported in <xref ref-type="bibr" rid="bib1.bibx57" id="text.73"/>. Since the unperturbed system has an equilibrium near the upper fold point, the motion is stagnant near the fold point. This provides favorable conditions for observing SPS. The state jumps from the upper branch of the critical manifold to its lower branch often occur after an upward jump induced by noise. These upward jumps resemble the rebound events prior to DO cooling transitions. The overall phenomenon is the same in the self-sustained oscillation regime of the FHN model, as long as the equilibrium is located near the upper fold point of the critical manifold (<inline-formula><mml:math id="M147" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>≃</mml:mo><mml:mn mathvariant="normal">0.25</mml:mn></mml:mrow></mml:math></inline-formula>).</p></list-item><list-item>
      <p id="d1e2489"><italic>Hopf bifurcation mechanism</italic>. In contrast to the fold bifurcation, the Hopf bifurcation manifests oscillatory instability <xref ref-type="bibr" rid="bib1.bibx79" id="paren.74"/>. In several ocean box models, the thermohaline circulation is destabilized via a Hopf bifurcation <xref ref-type="bibr" rid="bib1.bibx2 bib1.bibx53 bib1.bibx1 bib1.bibx75" id="paren.75"/>. It is also considered a potential generating mechanism of DO oscillations in a low-order coupled climate model <xref ref-type="bibr" rid="bib1.bibx83" id="paren.76"/> and in a comprehensive climate model <xref ref-type="bibr" rid="bib1.bibx64" id="paren.77"/>. Assume that the parameter <inline-formula><mml:math id="M148" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> decreases slowly in the FHN-type model (Fig. 5d and e). The underlying dynamics changes from the stable equilibrium to the limit-cycle oscillations at the Hopf bifurcation point <inline-formula><mml:math id="M149" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>Hopf</mml:mtext></mml:msub><mml:mo>≡</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>/</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>/</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula> (Appendix B). If stochastic forcing is added to the system, noise-induced small oscillations can appear prior to the onset of the limit-cycle oscillations (Fig. 5d and e). The precursor<?pagebreak page693?> oscillations resemble rebound events, while their shape depends on the noise as well as the change rate of <inline-formula><mml:math id="M150" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula>. Again SPS can be observed near the Hopf bifurcation point (Fig. S30) <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx57 bib1.bibx11" id="paren.78"/>. The small oscillations prior to downward transitions, like the DO rebound events, do not appear if the system goes deeply into the self-sustained oscillation regime away from the Hopf bifurcation point (<inline-formula><mml:math id="M151" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>&lt;</mml:mo><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>Hopf</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">0.245</mml:mn></mml:mrow></mml:math></inline-formula> in Fig. 5d).</p></list-item><list-item>
      <p id="d1e2589"><italic>Mixed-mode oscillation mechanism</italic>. Mixed-mode oscillations (MMOs) are periodic oscillations consisting of small- and large-amplitude oscillations <xref ref-type="bibr" rid="bib1.bibx43 bib1.bibx28 bib1.bibx6" id="paren.79"/>. They often arise in systems with one fast variable and two slow variables <xref ref-type="bibr" rid="bib1.bibx28" id="paren.80"/>. In this regard, we can extend the above FHN-type model to exhibit MMOs, for example, as follows: <inline-formula><mml:math id="M152" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M153" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>y</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi>k</mml:mi><mml:mo>(</mml:mo><mml:mi>z</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M154" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>z</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>z</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>z</mml:mi><mml:mo>+</mml:mo><mml:mi>k</mml:mi><mml:mo>(</mml:mo><mml:mi>y</mml:mi><mml:mo>-</mml:mo><mml:mi>z</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M155" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> is another slow variable with timescale <inline-formula><mml:math id="M156" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>z</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M157" display="inline"><mml:mrow><mml:mo>≫</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and <inline-formula><mml:math id="M158" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula> is the diffusive-coupling constant between slow variables. We interpret <inline-formula><mml:math id="M159" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> as the surface salinity in the northern North Atlantic convection region that directly affects the AMOC strength <inline-formula><mml:math id="M160" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> again and <inline-formula><mml:math id="M161" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> as the surface salinity outside the convection region that affects the surface salinity <inline-formula><mml:math id="M162" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> in the convection region via mixing. This extended FHN-type model is introduced here only to demonstrate that MMOs may appear in an FHN-type model with a minimal dimensional extension. For certain parameter settings (Appendix B), the system has an unstable equilibrium <inline-formula><mml:math id="M163" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>,</mml:mo><mml:mi>z</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> of saddle-focus type, with one stable direction with a negative real eigenvalue and a two-dimensional unstable manifold with complex eigenvalues with a positive real part. The slow–fast oscillations occur along the critical manifold <inline-formula><mml:math id="M164" display="inline"><mml:mrow><mml:mo mathvariant="italic">{</mml:mo><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>,</mml:mo><mml:mi>z</mml:mi><mml:mo>)</mml:mo><mml:mo>∈</mml:mo><mml:msup><mml:mi mathvariant="double-struck">R</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mo>|</mml:mo><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo mathvariant="italic">}</mml:mo></mml:mrow></mml:math></inline-formula> (Fig. 5f and g). However, due to the saddle-focus equilibrium on the critical manifold, the trajectory is attracted toward the saddle from the direction of the stable manifold (black segment) and repelled from it in a spiralling fashion. The striking point is the systematic occurrence of small-amplitude oscillations prior to the abrupt transition, which also resemble the rebound events prior to the DO cooling transitions. A more realistic time series is obtained if an observation noise is added on <inline-formula><mml:math id="M165" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>  (Fig. S31). Then SPS can be stably observed near the fold point of the critical manifold.</p></list-item></list> Based on the four types of simple mathematical models, we have proposed four possible dynamical mechanisms for the DO cooling transitions that can manifest statistical precursor signals (SPS): (1) strict CSD due to the approaching of a fold bifurcation; (2) CSD in a wider sense, in stochastic slow–fast oscillations; (3) noise-induced oscillations prior to Hopf bifurcations; or (4) the signal of mixed-mode oscillations. While the details of these mechanisms are different, they are all related to the fold points of the equilibrium curve or the critical manifolds. As a result, the SPS can be detected by the conventional CSD indicators.</p>
      <p id="d1e2942">Mechanisms (2), (3), and (4) can generate behavior resembling the rebound events, leading to increases in the classic CSD indicators. In the toy models, rebound event-like behavior is generated when the trajectory passes by an equilibrium point with marginal stability (i.e., the equilibrium has neither strong stability leading to a permanent state nor strong instability leading to short interstadials) (Fig. 5b–g). In this case, the duration of the modeled interstadial is relatively long in relation to the marginal stability. By contrast, the absence of equilibrium or the presence of a strongly unstable equilibrium near the fold point of the critical manifold leads to brief interstadials without a rebound event and consequently a lack of SPS. This provides a possible explanation of why the rebound events and the robust SPS are simultaneously observed for long interstadials but not for short interstadials.</p>
      <p id="d1e2945">Another possible explanation for the lack of SPS for short interstadials is the following. The common assumption underlying CSD theory is that the parameter change is much slower than the system's relaxation time, and the latter is much slower than the correlation time of the noise <xref ref-type="bibr" rid="bib1.bibx82 bib1.bibx4" id="paren.81"/>. If this assumption is violated, it is difficult to detect CSD-based SPS <xref ref-type="bibr" rid="bib1.bibx23 bib1.bibx84" id="paren.82"/>. Consider the fold-bifurcation-induced tipping in the Stommel-type model (1), for example (Fig. 6). If the change in the parameter <inline-formula><mml:math id="M166" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> is faster than the system's relaxation time toward the moving stable equilibrium, it is unlikely to detect significant CSD-based SPS (Fig. 6b) because the trajectory evolves systematically away from the equilibrium and thus cannot feel the flattening of the potential around the equilibrium even at the true bifurcation point.</p>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <label>5</label><title>Summary and discussion</title>
      <p id="d1e2970">In this study we have explored statistical precursor signals (SPSs) and significant increases in critical slowing down (CSD) indicators (variance and lag-1 autocorrelation), for Dansgaard–Oeschger (DO) cooling transitions following interstadials, using six Greenland ice core records. Among the 12 interstadials longer than 1000 years, we find at least one robust SPS for eight interstadials longer than roughly 1500 years (GI-25, GI-23.1, GI-21.1, GI-20, GI-19.2, GI-14, GI-12, and GI-8) and multiple robust SPSs for six of them (GI-25, GI-23.1, GI-21.1, GI-14, GI-12, and GI-8) (Fig. 4d). Robust SPSs are, however, not observed for interstadials shorter than roughly 1500 years. One might link the increase in the proxy variance with the tendency of larger climatic fluctuations in colder climates <xref ref-type="bibr" rid="bib1.bibx31" id="paren.83"/>, but the increases in the lag-1 autocorrelation cannot generally be<?pagebreak page694?> explained by it. The analysis removing the rebound events from the data shows that the rebound events prior to the cooling transitions are important in producing the SPS.</p>
      <p id="d1e2976">We have proposed four different dynamical mechanisms to explain the observed SPSs: (1) strict CSD due to the approaching of a fold bifurcation; (2) CSD in a wider sense, in stochastic slow–fast oscillations; (3) noise-induced oscillations prior to Hopf bifurcations; or (4) the signal of mixed-mode oscillations. In the last three mechanisms, oscillations such as the rebound events can systematically arise prior to the abrupt cooling transitions. These precursor oscillations are due to marginally (un)stable equilibria on the critical manifolds that cause a long-lived quasi-stable state (like long interstadials). This can explain why rebound events and SPSs are simultaneously observed only for long interstadials and are not observed for short ones. While the SPSs for bifurcation-induced tipping events (mechanisms 1 and 3) are established, detailed properties of SPSs for the stochastic slow–fast oscillations of the excitable system (mechanism 2) and for the mixed-mode oscillations (mechanism 4) remain to be elucidated.</p>
      <p id="d1e2979">We should mention the assumptions made in this study as well as alternative scenarios for the DO cooling transitions. First, the four dynamical mechanisms assume slow changes in parameters or slow variables which cause bifurcations in the fast subsystem. On the other hand, the rate-induced tipping mechanism has also been invoked for a possible AMOC collapse, where the rate of change of the external forcing (e.g., freshwater flux or atmospheric CO<inline-formula><mml:math id="M167" display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration) determines the future AMOC state <xref ref-type="bibr" rid="bib1.bibx2 bib1.bibx51 bib1.bibx70" id="paren.84"/>. The lack of observed SPSs for the interstadials less than roughly <inline-formula><mml:math id="M168" display="inline"><mml:mn mathvariant="normal">1500</mml:mn></mml:math></inline-formula> years indicates a rate-dependent aspect of the DO cooling transitions. However, a comprehensive investigation of DO cooling transitions from the viewpoint of rate-induced tipping is beyond the scope of this work. Second, a recent study using an ocean general circulation model shows that a rebound-event-type behavior of AMOC is caused by a behavior called “the intermediate tipping”, due to multiple stable ocean circulation states that exist near but prior to the tipping point leading to a significant AMOC weakening <xref ref-type="bibr" rid="bib1.bibx52" id="paren.85"/>. The intermediate tipping mechanism for rebound events is different from the possible low-dimensional dynamical mechanisms proposed in this study. Further studies are needed to elucidate the dynamical as well as physical origin of DO coolings and associated rebound events.</p>
      <p id="d1e3004">We have shown that past abrupt DO cooling transitions in the North Atlantic region can be anticipated based on classic CSD indicators if they are preceded by long interstadials. However, it is found to be difficult to anticipate DO cooling events, at least from the 20-year-resolution ice core Greenland records, if they occur after a short interstadial. If the DO coolings transitions are actually associated with AMOC weakening (see the “Introduction”), our results may have an implication for the predicted weakening of the AMOC and its possible collapse in the future: the prediction with CSD indicators could be more difficult if the forcing changes fast. There is, however, a caveat to this implication because the past DO cooling transitions are different from the presently inferred AMOC changes. The time resolution (mainly <inline-formula><mml:math id="M169" display="inline"><mml:mn mathvariant="normal">20</mml:mn></mml:math></inline-formula> years and additionally <inline-formula><mml:math id="M170" display="inline"><mml:mn mathvariant="normal">5</mml:mn></mml:math></inline-formula> years) and the length (mainly <inline-formula><mml:math id="M171" display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">1000</mml:mn></mml:mrow></mml:math></inline-formula> years and additionally <inline-formula><mml:math id="M172" display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">300</mml:mn></mml:mrow></mml:math></inline-formula> years) of the interstadial segment data used in this study are coarser and mostly longer than the annual data used for analyzing AMOC fingerprints during the industrial period <xref ref-type="bibr" rid="bib1.bibx8 bib1.bibx5 bib1.bibx29" id="paren.86"/> and the last millennium <xref ref-type="bibr" rid="bib1.bibx59" id="paren.87"/>. More crucially, the revealed predictability of past DO cooling events does not necessarily imply predictability of a potential future AMOC collapse since the recent AMOC weakening, possibly driven by global warming but potentially also part of natural variability, is mechanistically very different from past AMOC weakening in the glacial period.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <?xmltex \currentcnt{A}?><label>Appendix A</label><title>Probability of observing robust precursor signals</title>
      <p id="d1e3059">The statistical significance of precursor signals of critical transitions, in terms of positive trends of CSD indicators, is assessed by hypothesis testing  <xref ref-type="bibr" rid="bib1.bibx80 bib1.bibx26 bib1.bibx73 bib1.bibx7" id="paren.88"/>. We consider a stationary stochastic process with preserved variance and autocorrelation as the null model. The <inline-formula><mml:math id="M173" display="inline"><mml:mi>n</mml:mi></mml:math></inline-formula> surrogate data are prepared from the original residual data series by the phase-randomization method, thus preserving the variance and autocorrelation function of the original time series via the Wiener–Khinchin theorem. Here we take <inline-formula><mml:math id="M174" display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1000</mml:mn></mml:mrow></mml:math></inline-formula>. The linear trend (<inline-formula><mml:math id="M175" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>) of the CSD indicator for the original residual time series and the linear trends (<inline-formula><mml:math id="M176" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) of CSD indicators for the surrogate data are calculated. We consider the precursor signal of the original series statistically significant at the 5 % level if the probability of <inline-formula><mml:math id="M177" display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub><mml:mo>&gt;</mml:mo><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">o</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M178" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula> value) is less than 0.05. Thus, if the original data are already a stationary stochastic process (exhibiting no CSD), one should expect spuriously significant results at a probability of <inline-formula><mml:math id="M179" display="inline"><mml:mn mathvariant="normal">0.05</mml:mn></mml:math></inline-formula> by definition. In principle this is independent of the smoothing span <inline-formula><mml:math id="M180" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> as well as the rolling window size <inline-formula><mml:math id="M181" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> used for calculating CSD indicators. We consider a precursor signal robust if we find significant cases (<inline-formula><mml:math id="M182" display="inline"><mml:mrow><mml:mi>p</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>) for more than half (<inline-formula><mml:math id="M183" display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">15</mml:mn></mml:mrow></mml:math></inline-formula>) of 30 combinations of <inline-formula><mml:math id="M184" display="inline"><mml:mi mathvariant="italic">α</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M185" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula>. Then the probability of observing a robust precursor signal can be shown to be 0.05. In order to check this numerically, we generate 5000 surrogates of the original <inline-formula><mml:math id="M186" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">18</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>O series of interstadial GI-25 and calculate the probability of finding robust precursor signals. The resulting fractions are 0.041 for the variance and 0.039 for the lag-1 autocorrelation, which are close to 0.05. For the case of GI-12, we obtain 0.038 for the variance and 0.047 for the lag-1 autocorrelation, again close to 0.05. These results support the<?pagebreak page695?> probability of observing a robust precursor signal being 5 % if the data are stationary stochastic processes.</p>
</app>

<app id="App1.Ch1.S2">
  <?xmltex \currentcnt{B}?><label>Appendix B</label><title>Details of conceptual models used in Fig. 5</title>
      <p id="d1e3209">Here we describe specific settings for four conceptual models representing different candidate mechanisms for the DO cooling transitions. Unless otherwise mentioned, the stochastic differential equations below are solved with the Euler–Maruyama method with a step size of <inline-formula><mml:math id="M187" display="inline"><mml:mrow><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. <list list-type="order"><list-item>
      <p id="d1e3228">The bistability of the AMOC strength <inline-formula><mml:math id="M188" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> can be conceptually modeled by a double-fold bifurcation model: <inline-formula><mml:math id="M189" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M190" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> has two fold points; here for <inline-formula><mml:math id="M191" display="inline"><mml:mi>f</mml:mi></mml:math></inline-formula> one can use either <inline-formula><mml:math id="M192" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:msup><mml:mi>x</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> or <inline-formula><mml:math id="M193" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. We take the quadratic function <inline-formula><mml:math id="M194" display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> that arises in the <xref ref-type="bibr" rid="bib1.bibx78" id="text.89"/> model. <inline-formula><mml:math id="M195" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> represents a forcing parameter on the AMOC strength <inline-formula><mml:math id="M196" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>, e.g., salinity forcing on the North Atlantic (i.e., negative freshwater forcing). <inline-formula><mml:math id="M197" display="inline"><mml:mrow><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is white Gaussian noise, e.g., freshwater perturbations or weather forcing. In Fig. 5a, we set <inline-formula><mml:math id="M198" display="inline"><mml:mrow><mml:msqrt><mml:mrow><mml:mo>〈</mml:mo><mml:msup><mml:mi mathvariant="italic">ξ</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>〉</mml:mo></mml:mrow></mml:msqrt><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.03</mml:mn></mml:mrow></mml:math></inline-formula>, and the initial condition is taken at <inline-formula><mml:math id="M199" display="inline"><mml:mrow><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1.1</mml:mn></mml:mrow></mml:math></inline-formula>, near the upper stable fixed point of the unperturbed system. The parameter <inline-formula><mml:math id="M200" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> is then slowly decreased from <inline-formula><mml:math id="M201" display="inline"><mml:mn mathvariant="normal">0.1</mml:mn></mml:math></inline-formula> to <inline-formula><mml:math id="M202" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.4</mml:mn></mml:mrow></mml:math></inline-formula> over the period from <inline-formula><mml:math id="M203" display="inline"><mml:mrow><mml:mi>t</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:math></inline-formula> to <inline-formula><mml:math id="M204" display="inline"><mml:mrow><mml:mi>t</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">500</mml:mn></mml:mrow></mml:math></inline-formula>, to trigger the bifurcation-induced transition.</p></list-item><list-item>
      <p id="d1e3508">The FitzHugh–Nagumo-type (FHN-type) system is a prototypical model of slow–fast oscillators <xref ref-type="bibr" rid="bib1.bibx34 bib1.bibx62" id="paren.90"/> and often invoked for conceptual models of DO oscillations <xref ref-type="bibr" rid="bib1.bibx68 bib1.bibx46 bib1.bibx71 bib1.bibx61 bib1.bibx50 bib1.bibx69 bib1.bibx86" id="paren.91"/>. The FHN-type model subjected to dynamical noise can be obtained by introducing a slow variable <inline-formula><mml:math id="M205" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> into the fold bifurcation model: <inline-formula><mml:math id="M206" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi mathvariant="italic">ξ</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M207" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>y</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M208" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M209" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> are timescale parameters (<inline-formula><mml:math id="M210" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>≪</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>). Following the salt-oscillator hypothesis to explain DO cycles <xref ref-type="bibr" rid="bib1.bibx85" id="paren.92"/>, we may interpret <inline-formula><mml:math id="M211" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> as the salinity in the polar halocline surface mixed layer, which decreases (increases) when the AMOC is strong (weak). In the case of Fig. 5b and c, we set <inline-formula><mml:math id="M212" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.26</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M213" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.01</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M214" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M215" display="inline"><mml:mrow><mml:msqrt><mml:mrow><mml:mo>〈</mml:mo><mml:msup><mml:mi mathvariant="italic">ξ</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>〉</mml:mo></mml:mrow></mml:msqrt><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.3</mml:mn></mml:mrow></mml:math></inline-formula>, and the initial condition is taken at <inline-formula><mml:math id="M216" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.2</mml:mn><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.45</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. The <inline-formula><mml:math id="M217" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> nullcline (critical manifold) of the unperturbed system is <inline-formula><mml:math id="M218" display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mi mathvariant="italic">μ</mml:mi></mml:mrow></mml:math></inline-formula> (Fig. 5c, green) and the <inline-formula><mml:math id="M219" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> nullcline is <inline-formula><mml:math id="M220" display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>=</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi></mml:mrow></mml:math></inline-formula> (Fig. 5c, magenta dashed). The intersection of the <inline-formula><mml:math id="M221" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>- and <inline-formula><mml:math id="M222" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> nullclines is the equilibrium point of the unperturbed system <inline-formula><mml:math id="M223" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, which is near the fold point of the critical manifold in this parameter setting.</p></list-item><list-item>
      <p id="d1e3878">To demonstrate the Hopf bifurcation mechanism in Fig. 5d and e, the same stochastic FHN-type model is used with <inline-formula><mml:math id="M224" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.01</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M225" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M226" display="inline"><mml:mrow><mml:msqrt><mml:mrow><mml:mo>〈</mml:mo><mml:msup><mml:mi mathvariant="italic">ξ</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>〉</mml:mo></mml:mrow></mml:msqrt><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.05</mml:mn></mml:mrow></mml:math></inline-formula>, but here <inline-formula><mml:math id="M227" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula> is gradually decreased from <inline-formula><mml:math id="M228" display="inline"><mml:mn mathvariant="normal">0.3</mml:mn></mml:math></inline-formula> to <inline-formula><mml:math id="M229" display="inline"><mml:mn mathvariant="normal">0.2</mml:mn></mml:math></inline-formula>, over a period of 5 time units. For <inline-formula><mml:math id="M230" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.2</mml:mn><mml:mo>&lt;</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>&lt;</mml:mo><mml:mn mathvariant="normal">0.3</mml:mn></mml:mrow></mml:math></inline-formula>, the system has a unique equilibrium point at <inline-formula><mml:math id="M231" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>. The Hopf bifurcation of an equilibrium occurs if the complex eigenvalues of the Jacobian matrix at the equilibrium pass the imaginary axis <xref ref-type="bibr" rid="bib1.bibx79" id="paren.93"/>. The eigenvalues of the Jacobian matrix at this equilibrium are <inline-formula><mml:math id="M232" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">λ</mml:mi><mml:mo>±</mml:mo></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">2</mml:mn></mml:mfrac></mml:mstyle><mml:mo mathvariant="italic">{</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>±</mml:mo><mml:msqrt><mml:mrow><mml:mo>(</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:mn mathvariant="normal">8</mml:mn><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:msqrt><mml:mo mathvariant="italic">}</mml:mo></mml:mrow></mml:math></inline-formula>. These eigenvalues <inline-formula><mml:math id="M233" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">λ</mml:mi><mml:mo>±</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula> are complex conjugates for <inline-formula><mml:math id="M234" display="inline"><mml:mrow><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">4</mml:mn></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>+</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:msqrt><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:msqrt><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>&lt;</mml:mo><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>&lt;</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">4</mml:mn></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>+</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>+</mml:mo><mml:mn mathvariant="normal">2</mml:mn><mml:msqrt><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:msqrt><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>. In this range of <inline-formula><mml:math id="M235" display="inline"><mml:mi mathvariant="italic">μ</mml:mi></mml:math></inline-formula>, the real part of <inline-formula><mml:math id="M236" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">λ</mml:mi><mml:mo>±</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula> changes from negative to positive at the Hopf bifurcation point: <inline-formula><mml:math id="M237" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>Hopf</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">4</mml:mn></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>.</mml:mo></mml:mrow></mml:math></inline-formula> For <inline-formula><mml:math id="M238" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>/</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.01</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M239" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">μ</mml:mi><mml:mtext>Hopf</mml:mtext></mml:msub><mml:mo>≈</mml:mo><mml:mn mathvariant="normal">0.245</mml:mn><mml:mo>.</mml:mo></mml:mrow></mml:math></inline-formula> The initial condition is taken at the origin.</p></list-item><list-item>
      <p id="d1e4350">The mixed-mode oscillation model is obtained if the FHN-type model is extended to have multiple interacting slow variables. For example, <inline-formula><mml:math id="M240" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>x</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:mi>x</mml:mi><mml:mo>|</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi mathvariant="italic">μ</mml:mi></mml:mrow></mml:math></inline-formula>), <inline-formula><mml:math id="M241" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>y</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>+</mml:mo><mml:mi>k</mml:mi><mml:mo>(</mml:mo><mml:mi>z</mml:mi><mml:mo>-</mml:mo><mml:mi>y</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M242" display="inline"><mml:mrow><mml:mover accent="true"><mml:mi>z</mml:mi><mml:mo mathvariant="normal">˙</mml:mo></mml:mover><mml:mo>=</mml:mo><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>z</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>(</mml:mo><mml:mo>-</mml:mo><mml:mi>x</mml:mi><mml:mo>-</mml:mo><mml:mi>z</mml:mi><mml:mo>+</mml:mo><mml:mi>k</mml:mi><mml:mo>(</mml:mo><mml:mi>y</mml:mi><mml:mo>-</mml:mo><mml:mi>z</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, where <inline-formula><mml:math id="M243" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> is another slow variable with timescale <inline-formula><mml:math id="M244" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>z</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M245" display="inline"><mml:mrow><mml:mo>≫</mml:mo><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and <inline-formula><mml:math id="M246" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula> is the diffusive coupling constant between slow variables. We interpret <inline-formula><mml:math id="M247" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> as the surface salinity in the northern North Atlantic convection region, which directly affects the AMOC strength <inline-formula><mml:math id="M248" display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M249" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> as the surface salinity outside the convection region that affects the surface salinity <inline-formula><mml:math id="M250" display="inline"><mml:mi>y</mml:mi></mml:math></inline-formula> in the convection region via mixing. We set <inline-formula><mml:math id="M251" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>x</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.02</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M252" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>y</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M253" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>z</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">4</mml:mn></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M254" display="inline"><mml:mrow><mml:mi mathvariant="italic">μ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.225</mml:mn></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M255" display="inline"><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.8</mml:mn></mml:mrow></mml:math></inline-formula>. This system has an unstable equilibrium <inline-formula><mml:math id="M256" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>,</mml:mo><mml:mi>z</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:msqrt><mml:mi mathvariant="italic">μ</mml:mi></mml:msqrt><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> of saddle-focus type, with one stable direction with a negative real eigenvalue of <inline-formula><mml:math id="M257" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.67</mml:mn></mml:mrow></mml:math></inline-formula> and a two-dimensional unstable manifold with two complex conjugate eigenvalues with a positive real part of <inline-formula><mml:math id="M258" display="inline"><mml:mrow><mml:mn mathvariant="normal">0.94</mml:mn><mml:mo>±</mml:mo><mml:mn mathvariant="normal">4.7</mml:mn><mml:mi>i</mml:mi></mml:mrow></mml:math></inline-formula>. The initial condition is taken at <inline-formula><mml:math id="M259" display="inline"><mml:mrow><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>,</mml:mo><mml:mi>y</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>,</mml:mo><mml:mi>z</mml:mi><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>)</mml:mo><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0.5</mml:mn><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.5</mml:mn><mml:mo>,</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">0.5</mml:mn><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>.</p></list-item></list></p>
</app>
  </app-group><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d1e4773">The Greenland ice core records used in this study can be obtained from <uri>https://www.iceandclimate.nbi.ku.dk/data/</uri> <xref ref-type="bibr" rid="bib1.bibx21" id="paren.94"/>. The R codes used in this study are available from <ext-link xlink:href="https://doi.org/10.5281/zenodo.10841655" ext-link-type="DOI">10.5281/zenodo.10841655</ext-link> <xref ref-type="bibr" rid="bib1.bibx60" id="paren.95"/>.</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d1e4788">The supplement related to this article is available online at: <inline-supplementary-material xlink:href="https://doi.org/10.5194/cp-20-683-2024-supplement" xlink:title="pdf">https://doi.org/10.5194/cp-20-683-2024-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d1e4797">TM conceived the study and conducted the analyses with contributions from NB. Both authors discussed<?pagebreak page696?> and interpreted the results. TM wrote the manuscript with contributions from NB.</p>
  </notes><?xmltex \hack{\newpage}?><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d1e4804">The contact author has declared that neither of the authors has any competing interests.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d1e4810">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. While Copernicus Publications makes every effort to include appropriate place names, the final responsibility lies with the authors.</p>
  </notes><ack><title>Acknowledgements</title><p id="d1e4816">The authors thank Keno Riechers and Maya Ben-Yami for their helpful comments.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d1e4821">The authors acknowledge funding by the Volkswagen Foundation. This is TiPES contribution #243; The TiPES (“Tipping Points in the Earth System”) project has received funding from the European Union's Horizon 2020 research and innovation programme under grant agreement no. 820970. Niklas Boers acknowledges further funding by the European Union's Horizon 2020 research and innovation programme under the Marie Sklodowska-Curie grant agreement no. 956170, as well as from the Federal Ministry of Education and Research under grant no. 01LS2001A.</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d1e4828">This paper was edited by Bjørg Risebrobakken and reviewed by three anonymous referees.</p>
  </notes><ref-list>
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