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  <front>
    <journal-meta>
<journal-id journal-id-type="publisher">CP</journal-id>
<journal-title-group>
<journal-title>Climate of the Past</journal-title>
<abbrev-journal-title abbrev-type="publisher">CP</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Clim. Past</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">1814-9332</issn>
<publisher><publisher-name>Copernicus Publications</publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>

    <article-meta>
      <article-id pub-id-type="doi">10.5194/cp-12-807-2016</article-id><title-group><article-title>Model simulations of early westward flow across<?xmltex \hack{\newline}?> the Tasman Gateway during the early Eocene</article-title>
      </title-group><?xmltex \runningtitle{Model simulations of early westward flow across the Tasman Gateway}?><?xmltex \runningauthor{W.~P.~Sijp et al.}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Sijp</surname><given-names>Willem P.</given-names></name>
          <email>w.sijp@unsw.edu.au</email>
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>von der Heydt</surname><given-names>Anna S.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-5557-3282</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3">
          <name><surname>Bijl</surname><given-names>Peter K.</given-names></name>
          
        </contrib>
        <aff id="aff1"><label>1</label><institution>ARC Centre of Excellence for Climate System Science, University of New South
Wales, Sydney, NSW 2052, Australia</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Institute for Marine and Atmospheric Research, Centre for Extreme Matter and Emergent Phenomena, Utrecht University, Princetonplein 5, 3584 CC Utrecht, the Netherlands</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Marine Palynology and Paleoceanography, Laboratory of Palaeobotany and Palynology, Department of Earth Sciences, Faculty of Geoscience, Utrecht University, P.O. Box 80115, 3508 TC Utrecht, the Netherlands</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Willem P. Sijp (w.sijp@unsw.edu.au)</corresp></author-notes><pub-date><day>5</day><month>April</month><year>2016</year></pub-date>
      
      <volume>12</volume>
      <issue>4</issue>
      <fpage>807</fpage><lpage>817</lpage>
      <history>
        <date date-type="received"><day>10</day><month>July</month><year>2015</year></date>
           <date date-type="rev-request"><day>29</day><month>October</month><year>2015</year></date>
           <date date-type="rev-recd"><day>17</day><month>February</month><year>2016</year></date>
           <date date-type="accepted"><day>6</day><month>March</month><year>2016</year></date>
      </history>
      <permissions>
<license license-type="open-access">
<license-p>This work is licensed under a Creative Commons Attribution 3.0 Unported License. To view a copy of this license, visit <ext-link ext-link-type="uri" xlink:href="http://creativecommons.org/licenses/by/3.0/">http://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions><self-uri xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016.html">This article is available from https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016.html</self-uri>
<self-uri xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016.pdf">The full text article is available as a PDF file from https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016.pdf</self-uri>


      <abstract>
    <p>The timing and role in ocean circulation and climate of the
opening of Southern Ocean gateways is as yet elusive. Recent
micropalaeontological studies suggest the onset of westward throughflow of
surface waters from the SW Pacific into the Australo-Antarctic Gulf through a
southern shallow opening of the Tasman Gateway from 49–50 Ma onwards, a
direction that is counter to the present-day eastward-flowing Antarctic
Circumpolar Current. Here, we present the first model results specific to the
early-to-middle Eocene where, in agreement with the field evidence, southerly
shallow opening of the Tasman Gateway indeed causes a westward flow across
the Tasman Gateway. As a result, modelled estimates of dinoflagellate
biogeography are in agreement with the recent findings. Crucially, in this
situation where Australia is still situated far south and almost attached to
Antarctica, the Drake Passage must be sufficiently restricted to allow the
prevailing easterly wind pattern to set up this southerly restricted westward
flow. In contrast, an open Drake Passage, up to 517 m deep, leads to an
eastward flow, even when the Tasman Gateway and the Australo-Antarctic gulf
are entirely contained within the latitudes of easterly wind.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <title>Introduction</title>
      <p>The different positions of large continents in the Southern Hemisphere
influenced the oceanography of the Eocene and thus the distribution of
heat. The position of Australia and South America in the Eocene was much
closer to Antarctica than today <xref ref-type="bibr" rid="bib1.bibx8" id="paren.1"><named-content content-type="pre">see, e.g.,</named-content></xref>. This arguably
prevented circumpolar surface water flow
<xref ref-type="bibr" rid="bib1.bibx22 bib1.bibx5 bib1.bibx6" id="paren.2"/>, i.e. the Antarctic Circumpolar
Current (ACC), that thermally isolates Antarctica today
<xref ref-type="bibr" rid="bib1.bibx43 bib1.bibx36" id="paren.3"/>. Instead, it was thought that low-latitude
currents bathed Antarctic coastlines. The subsequent opening of a circumpolar
passage, through deepening of the Drake Passage (DP) and Tasman Gateway (TG),
would then allow the ACC to develop, leading to the isolation of Antarctica and
progressive cooling, ultimately leading to full glaciation of Antarctica
<xref ref-type="bibr" rid="bib1.bibx23" id="paren.4"/>. Support for this hypothesis came when it was discovered
that the deep opening of these gateways appeared to be roughly coeval with
the onset of continental-scale Antarctic glaciation around the
Eocene–Oligocene transition <xref ref-type="bibr" rid="bib1.bibx2" id="paren.5"><named-content content-type="pre">EOT; about 34 million years ago,
Ma;</named-content></xref>.</p>
      <p>Objections to the gateway hypothesis came from two fields of research. First
of all, drill cores obtained from the Tasman Gateway region showed that
accelerated widening and deepening of the passage likely started 2 Myr prior
to the glaciation, indicating that a direct causal link between the two is
unlikely <xref ref-type="bibr" rid="bib1.bibx42" id="paren.6"/>. Unlike the quite precise estimates for the
opening of the Tasman Gateway, the timing of the opening of the Drake Passage is
debated and less well constrained
<xref ref-type="bibr" rid="bib1.bibx1 bib1.bibx26 bib1.bibx35 bib1.bibx33" id="paren.7"/>. Several
structural geologic investigations in the region have inferred crustal
stretching from the early Eocene onwards
<xref ref-type="bibr" rid="bib1.bibx25 bib1.bibx17" id="paren.8"/>, but it remains unclear whether the
vertical displacement of that tectonism caused a shallow passageway through
the Drake Passage. We cannot, however, rule out the possibility that the Drake
Passage was already open, but shallow, in the early Eocene.</p>
      <p>Secondly, numerical modelling studies with general circulation models (GCMs)
configured for the Eocene emerged that suggested an ocean circulation pattern
with clockwise circulating gyres <xref ref-type="bibr" rid="bib1.bibx41 bib1.bibx21 bib1.bibx22" id="paren.9"/>,
where western boundary currents near Antarctica (as part of a subpolar gyre)
still prevented low-latitude currents from reaching and warming Antarctica (a general
feature of subpolar gyres). Microfossil biogeography studies (notably, but
not exclusively, done on organic-walled dinoflagellate cysts) clearly support
the gyre configuration as suggested by numerical modelling experiments
<xref ref-type="bibr" rid="bib1.bibx22 bib1.bibx5" id="paren.10"/> rather than the strong influence of
low-latitude-derived ocean currents. It should be noted here that these
considerations of the gyre circulation alone are centred on the surface
circulation and do not include influences of thermohaline heat transport,
which has been shown to have the potential to significantly enhance oceanic
heat transport to Antarctica in a closed-gateway configuration
<xref ref-type="bibr" rid="bib1.bibx38 bib1.bibx39 bib1.bibx46" id="paren.11"/>.</p>
      <p>The existing proxy records for atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> during the Paleogene
<xref ref-type="bibr" rid="bib1.bibx31 bib1.bibx32 bib1.bibx3" id="paren.12"/> and ice sheet modelling
<xref ref-type="bibr" rid="bib1.bibx10 bib1.bibx11" id="paren.13"/> have steered opinion in the direction of
declining atmospheric greenhouse gas concentrations as the primary forcing
factor in explaining Eocene cooling in general and, ultimately, the onset of
continental-scale glaciation.</p>
      <p>Although gateway changes appear not to be the direct cause of EOT glaciation,
unlike the hypothesis first proposed by <xref ref-type="bibr" rid="bib1.bibx23" id="text.14"/> and <xref ref-type="bibr" rid="bib1.bibx24" id="text.15"/>, their role
in the general long-term Cenozoic cooling trend remains plausible
<xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx6 bib1.bibx40" id="paren.16"/> but poorly reconciled with data.
Indeed, glaciation (34 Ma) took place against the backdrop of a long and
gradual cooling trend in the Southern Ocean <xref ref-type="bibr" rid="bib1.bibx4" id="paren.17"/>, whereas
detailed geological reconstructions of the Tasman Gateway have, until
recently <xref ref-type="bibr" rid="bib1.bibx6" id="paren.18"/>, largely ignored the potential climatic effects
of earlier stages of its opening. Immediately preceding this slow climatic
deterioration is the time with the warmest global temperatures of the past 85
million years, the protracted greenhouse episode known as the Early Eocene
Climatic Optimum <xref ref-type="bibr" rid="bib1.bibx47" id="paren.19"><named-content content-type="pre">EECO; 52–50 Ma;</named-content></xref>. Whereas it is
generally believed that a decline in atmospheric greenhouse gas
concentrations terminated the EECO, compelling direct and unequivocal proxy
evidence for this is generally lacking.</p>
      <p>As a contrasting hypothesis, <xref ref-type="bibr" rid="bib1.bibx6" id="text.20"/> find evidence for the first
signs of throughflow of south-west (SW) Pacific surface waters into the
Australo-Antarctic gulf (AAG) across the Tasman Gateway coinciding with the
onset of regional surface water and continental Antarctic cooling of
2–4<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>. <xref ref-type="bibr" rid="bib1.bibx6" id="text.21"/> infer from the distinct difference in
dinocyst assemblages between the AAG and the SW Pacific that the Tasman
Gateway served as an effective barrier to surface water exchange before
50 Ma. From 49–50 Ma onwards, endemic dinocyst species originating in the
SW Pacific began to dominate sediments of the Antarctic margin of the AAG.
They propose that the timing is consistent with the drowning of continental
blocks in the southern part of the Tasman Gateway inferred from accelerated
rifting during the early-to-middle Eocene transition
<xref ref-type="bibr" rid="bib1.bibx9 bib1.bibx20" id="paren.22"><named-content content-type="pre">52–48 Ma;</named-content></xref>. Surface water throughflow of SW
Pacific surface waters into the AAG would bring the endemic Antarctic species
into the AAG. A crucial observation here is that the south Australian margin
remains isolated from the SW Pacific influence but remains exclusively
inhabited by cosmopolitan and low-latitude-derived dinocyst species.
Furthermore, the influence of surface water throughflow from the AAG into the
south-west Pacific remains restricted as well: no low-latitude-derived species
appear in the SW Pacific Ocean. Eastward flow through the Tasman Gateway
would be expected to bring the low-latitude species from the Australian margin
into the SW Pacific Ocean. This is contrary to their findings; therefore,
they hypothesize that the throughflow must have occurred to the south, perhaps within
the reach of the easterlies, allowing the throughflow of the taxa present in
the Ross Sea that consistently dominate the westward flow along Antarctica,
the Eocene Antarctic Counter Current.</p>
      <p>Organic biomarker proxy records for the paleotemperature of the sea surface
and for the air temperature were derived from the same sedimentary archives as the
oceanographic reconstructions. They show that the opening of the Tasmanian
Gateway coincided with surface water and air temperature cooling of several
degrees (2–4 <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C), with the Antarctic hinterland cooling the most
(4 <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C). Simultaneously, benthic foraminiferal oxygen isotope records
show the onset of gradual cooling as well. Although this study could not
prove causality between oceanographic changes and cooling, the closeness in
time is intriguing and requires a follow-up to prove causality.</p>
      <p>The uncertainty related to the role of gateways in long-term climate
evolution, as well as the increasing need for more detailed knowledge of
Cenozoic ocean current development in general, stimulates modelling studies on
the impact of gateway changes on ocean circulation through time. Here, we
will use a coupled climate model of intermediate complexity
<xref ref-type="bibr" rid="bib1.bibx39" id="paren.23"/> to numerically simulate the effect of an initial
opening of the Tasman Gateway at the early-to-middle Eocene transition. We
show that this can lead to the inferred patterns <xref ref-type="bibr" rid="bib1.bibx6" id="paren.24"/> of
dinoflagellate biogeography: a westward current emerges, but only when the
Drake Passage remains closed. Numerical modelling provides a physical basis to
explain micropalaeontological observations.</p>
</sec>
<sec id="Ch1.S2">
  <title>Model and experimental design</title>
      <p>We use a modified version of the intermediate-complexity coupled model
described in detail in <xref ref-type="bibr" rid="bib1.bibx45" id="text.25"/>, the so-called UVic (University of Victoria) model. The model
consists of an ocean general circulation model <xref ref-type="bibr" rid="bib1.bibx30" id="paren.26"><named-content content-type="pre">GFDL MOM, Geophysical Fluid Dynamics Laboratory Modular Ocean Model, Version
2.2;</named-content></xref> coupled to a simplified one-layer energy–moisture
balance model for the atmosphere and a dynamic–thermodynamic sea-ice model.
Air–sea heat and freshwater fluxes evolve freely in the model, while a
non-interactive wind field is employed for reasons of computational speed.
The turbulent kinetic energy scheme of <xref ref-type="bibr" rid="bib1.bibx7" id="text.27"/> based on
<xref ref-type="bibr" rid="bib1.bibx16" id="text.28"/> models vertical mixing due to wind and vertical velocity
shear. The model is identical to <xref ref-type="bibr" rid="bib1.bibx39" id="text.29"/>, with a modification
to the Eocene geography in the primary experiments where Australia is located
further south by 6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> in latitude and the Antarctic margin facing it
is also shifted south.</p>
      <p>The original geography will also be discussed as a secondary set of
experiments in the sensitivity study below, where both geography is altered and the Southern Hemisphere westerlies are shifted. Motivation for these
sensitivity studies of the true paleolatitude of the Tasman Gateway arises as
comparisons of Eocene palaeogeographies may vary by about 6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> in the SW
Pacific region dependent on which reference frame is used: a hotspot versus a
paleomagnetic reference frame <xref ref-type="bibr" rid="bib1.bibx44" id="paren.30"/>.</p>
      <p>We have run the model to equilibrium for a period of 9000 years in four configurations in which the Drake Passage is open to 517 m depth and in
simulations where it is closed; in each case the Tasman Gateway is
open, to a shallow depth of around 350 m, and closed. This choice is
informed by the expectation that the state of the Drake Passage exerts a
strong influence on the result of the opening of the Tasman Gateway: the
uncertainty surrounding the timing of the opening of the Drake Passage
<xref ref-type="bibr" rid="bib1.bibx25" id="paren.31"><named-content content-type="pre">see, e.g.,</named-content></xref> makes it necessary for us to consider an open and
a closed case.</p>
      <p>Limitations arise from the absence of a fully coupled ocean and atmosphere in the
UVic model. However, this model allows high enough spatial resolution to
represent the shallow, southern opening of the Tasmanian Gateway during the
early Eocene combined with long enough integration times to equilibrate the
deep ocean to examine a possible global temperature signal. Furthermore, it
provides the flexibility to conduct many simulations at once, each
representing different scenarios. To provide backup for the realistic set-up
of our model and cover both uncertainty in the location of the wind fields and their possible changes, our study prominently features experiments where
the winds have been shifted, precisely to test the robustness of the results and
cover the changes a fully coupled system might exhibit. Furthermore, a
simulation with an alternative wind field is explored in the Appendix.
Nonetheless, there could potentially be benefits to examining first-order
geostrophic feedbacks on the wind, an option that is present in the UVic
model.</p>
</sec>
<sec id="Ch1.S3">
  <title>Results</title>
      <p>The latitudinal section of the zonal velocity through the TG gap for the DP-closed case is shown in Fig. <xref ref-type="fig" rid="Ch1.F1"/>a and for the DP-open case in
Fig. <xref ref-type="fig" rid="Ch1.F2"/>a. In the simulation with the southern Tasman
Gateway open, a closed DP leads to a westward flow throughout the water
column near Antarctica and a shallower eastward flow to the north
(Fig. <xref ref-type="fig" rid="Ch1.F1"/>a). In contrast, the flow is eastward throughout
the gap when the DP is open (Fig. <xref ref-type="fig" rid="Ch1.F2"/>a). In this
simulation, the eastward flow is weak throughout most of the gap but strong
at its northern margin.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1"><caption><p>Meridional section of zonal velocity <inline-formula><mml:math display="inline"><mml:mi>u</mml:mi></mml:math></inline-formula> (cm s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) inside the
Tasman Gateway with the Drake Passage closed for <bold>(a)</bold> the standard
simulation, <bold>(b)</bold> the Southern Hemisphere westerlies shifted
6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> northward, and <bold>(c)</bold> Australia and the Antarctic margin in a more
northward position. All locations are well south of the latitudes of zero
wind stress curl. Positive values indicate eastward flow; negative values
indicate westward flow. </p></caption>
        <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f01.png"/>

      </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2"><caption><p>Meridional section of zonal velocity <inline-formula><mml:math display="inline"><mml:mi>u</mml:mi></mml:math></inline-formula> (cm s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) inside the
Tasman Gateway with the Drake Passage open for <bold>(a)</bold> the standard
simulation, <bold>(b)</bold> the Southern Hemisphere westerlies shifted
6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> northward, and <bold>(c)</bold> Australia and the Antarctic margin in
a more northward position. All locations are well south of the latitudes of
zero wind stress curl.</p></caption>
        <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f02.png"/>

      </fig>

      <p>To examine the sensitivity of these results to boundary conditions, we conducted two additional simulations where the core of the Southern
Hemisphere (SH) westerlies is shifted 6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> north, the TG is open, and
the DP is open and closed (thus yielding two simulations). For reference, we
show the zonal wind stress overlaid on the horizontal stream function in
Fig. <xref ref-type="fig" rid="Ch1.F3"/>: the average latitude of maximum westerly wind
stress (zero wind stress curl) is 5<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S in the standard case and
44<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S latitude in the wind-shifted case. This wind shifting
procedure is essentially equivalent to <xref ref-type="bibr" rid="bib1.bibx37" id="text.32"/>, and we refer to
this work for reference. We will refer to the original simulations without
the modification as the “standard” case. In this modified model
configuration, the flow pattern (Fig. <xref ref-type="fig" rid="Ch1.F1"/>b) is very
similar to the standard case with the DP closed (Fig. <xref ref-type="fig" rid="Ch1.F1"/>a).
Similarly, in the DP-open case, the flow pattern remains easterly throughout
the TG gap, again with weak flow through most of the gap, with the exception
of the northern margin (Fig. <xref ref-type="fig" rid="Ch1.F2"/>b). Note that, in
contrast to the corresponding standard simulation
(Fig. <xref ref-type="fig" rid="Ch1.F2"/>a), there is weak westward flow in a very narrow
band below 250 m depth; nonetheless, the general flow is overwhelmingly
eastward.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3"><caption><p>Ocean horizontal stream function (Sverdrup; 1 Sv is
<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mn>10</mml:mn><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:math></inline-formula> s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>), with the zonal wind stress (<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mn>10</mml:mn><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> Pa) overlaid.
The Drake Passage is closed for <bold>(a)</bold> the standard simulation,
<bold>(b)</bold> the Southern Hemisphere westerlies shifted north, and
<bold>(c)</bold> Australia and part of the Antarctic coast shifted north.</p></caption>
        <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f03.pdf"/>

      </fig>

      <p>In addition, we also conducted two simulations, again with the
TG open and the DP open and closed but now with altered geography. Here, the
Australian continent is shifted north by 6<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> latitude with respect to
our standard simulation, along with some northward shift of the Antarctic
margin of the AAG <xref ref-type="bibr" rid="bib1.bibx39" id="paren.33"><named-content content-type="pre">thus obtaining the original geography used
in</named-content></xref>. A southward-extending peninsula is also attached at the
location of Tasmania, in agreement with <xref ref-type="bibr" rid="bib1.bibx6" id="text.34"/>. Again, a similar
flow pattern emerges, where flow is eastward throughout the TG gap in the DP-open case (Fig. <xref ref-type="fig" rid="Ch1.F2"/>c), while there is a westward current
near Antarctica in the DP-closed case (Fig. <xref ref-type="fig" rid="Ch1.F1"/>c). We
conclude from our four additional simulations that our results are robust
with respect to the location of the major wind circulation patterns and
geography.</p>
      <p>From here on, we will focus on the standard simulations. First, we examine
the oceanic horizontal circulation. With the DP closed, the circulation of the
Southern Ocean is split into two subpolar clockwise gyres, the
Ross Sea gyre to the east of Australia and the Weddell gyre spanning
the (south) Indian and Atlantic oceans to the east of the DP. To the north lie
the subtropical gyres, where a super-gyre spans the Indian and Pacific
oceans. In agreement with the Sverdrup balance, eastward flow takes place at
the latitudes of positive wind stress curl (approximately minus the
latitudinal derivative of the wind stress shown in
Fig. <xref ref-type="fig" rid="Ch1.F3"/>). A proto-ACC is absent due to the closure of
the DP. Practically all eastward flow that passes Australia does so to the
north of the continent and is part of the southern branch of the subtropical
super-gyre.</p>
      <p>Eastward flow in the northern branches of the subpolar gyres generally
ends by returning south along the western margins of Australia and
South America when the DP is closed (Fig. <xref ref-type="fig" rid="Ch1.F1"/>a). A
discussion of the possibility of a linkage between the two subpolar gyres,
using additional simulations with another model, can be found in Appendix A.
Although the barotropic stream function shows no flow at the selected contour
interval, the velocity field indicates a westward flow in the Tasman Gateway
closely restricted to the Antarctic margin, naturally consistent with
Fig. <xref ref-type="fig" rid="Ch1.F1"/>a. This westward flow through the TG is
consistent with the generally negative wind stress curl at these latitudes.</p>
      <p>An open DP under otherwise early Eocene boundary conditions yields around
35 Sv flow through this gap, as read from the 5 Sv interval contours
immediately west of the DP (Fig <xref ref-type="fig" rid="Ch1.F4"/>a) and as determined by closer
analysis on the computer of the numerical model output (figure not shown).
The 35 Sv throughflow at the DP leads to eastward flow through the TG of around
8 Sv as determined from the model output file by computer analysis and shown
less precisely in Fig <xref ref-type="fig" rid="Ch1.F4"/>a. The remaining 27 Sv must pass north of
Australia. This total of 35 Sv of flow constitutes a circumpolar flow
component that is not part of the gyre structure and is reminiscent of the
ACC, despite the lack of latitudinal alignment of the gateways. We stress
that this flow is much weaker than today and not comparable, even
qualitatively. Nonetheless, we will refer to this current as the “proto-ACC”. As such, the uniformly eastward flow through the TG can be viewed as a
consequence of the flow through the DP gap that joins the eastward flow
along the northern branch of the Weddell gyre that must split to flow around
Australia, yielding a southern component. In addition, when opening the Drake
Passage, there is less necessity for significant westward return flow in the
subpolar gyres. Indeed, high-resolution ocean model simulations have shown
that closing the DP forces the ACC water to mainly turn southwards and join
the subpolar gyres (unpublished results). The circumpolar flow is in line
with idealized ocean modelling studies suggesting that strong circumpolar flow
is possible even without a fully unrestricted latitude band
<xref ref-type="bibr" rid="bib1.bibx29" id="paren.35"/>. This eastward flow occurs entirely within the
latitudes of the polar easterlies in the model
(Fig. <xref ref-type="fig" rid="Ch1.F3"/>) and within the latitudes of negative wind
stress curl (with a small exception in the wind-shifted case). This shows
that the direction of the flow is not only determined by the wind field but
also by the continental geometry.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4"><caption><p>Annual and vertical average of zonal velocity (cm s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>;
positive values indicate eastward flow) with ocean horizontal stream function
(Sverdrup; 1 Sv is <inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mn>10</mml:mn><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">3</mml:mn></mml:msup></mml:math></inline-formula> s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) overlaid. The Tasman Gateway is
open, with <bold>(a)</bold> the Drake Passage (DP) closed and <bold>(b)</bold> the DP
open (shallow at around 350 m depth). Contour intervals have been adjusted
in <bold>(a)</bold> to elucidate the flow through the TG.</p></caption>
        <?xmltex \igopts{width=239.00315pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f04.png"/>

      </fig>

<sec id="Ch1.S3.SSx1" specific-use="unnumbered">
  <title>Dinoflagellate biogeography and ocean circulation</title>
      <p>To examine links between Eocene ocean circulation and dinoflagellate
biogeography, we conduct several decadal timescale simulations, starting
from model equilibria, where a passive tracer (i.e. not interacting with the
ocean flow and representing small entities that do not move of their own
accord) is released at two locations, one south-east and one west of
Australia. We assume here that the current velocities of the surface water
dominate the active swim velocities of the dinoflagellates themselves. In any case, surface sediment analyses in the North Atlantic show a clear dominance of
specific Gulf Stream-loving dinocysts within the pathway of the Gulf Stream
<xref ref-type="bibr" rid="bib1.bibx48" id="paren.36"><named-content content-type="pre">e.g.</named-content></xref>, confirming the strong governance by
oceanography of biogeography of dinocyst species.</p>
      <p><?xmltex \hack{\newpage}?>Again, we perform two sets of simulations: a first set with the DP closed and
open or closed TG and a second set where the DP is open and the TG is open or closed.
The results of the tracer simulations are shown in
Figs. <xref ref-type="fig" rid="Ch1.F5"/> and <xref ref-type="fig" rid="Ch1.F6"/>, where the sites of the release of the tracer are marked by an
ellipse. We examine a snapshot taken 5 years after tracer release. The tracer
is interpreted as a fractional dinoflagellate concentration, and its
subsequent dispersal allows an estimate of expected dinoflagellate
biogeography based on ocean flow alone. The initial condition consists of the instantaneous insertion of a (rotationally symmetric)
Gaussian distribution of tracer in the horizontal direction of amplitude 1
(the tracer concentration is close to fraction 1, or 100 %, at the core
of the anomaly) and a horizontal standard deviation of several model grid
cells, allowing tracer concentration to approach zero away from the core and
similarly rapidly decreasing concentrations with depth according to a smaller
standard deviation appropriate to the vertical scale.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5"><caption><p>Dispersal of passive tracer representing dinoflagellates for the Drake
Passage closed and for the Tasman Gateway (TG) closed, originating west
<bold>(a)</bold> and east <bold>(b)</bold> of Australia, and for TG open, originating
west <bold>(c)</bold> and east <bold>(d)</bold> of Australia. Circles indicate the
release sites. When the TG is open, dinoflagellates released to the west of
Australia enter the Australo-Antarctic Gulf predominantly along the
Australian coast but remain west of the TG and do not enter the Ross Sea
<bold>(c)</bold>. In contrast, species released near the Ross Sea may enter the
gulf via a westward flow along the Antarctic coast <bold>(d)</bold>.
Concentrations are in percentages, where the initial localized concentration
upon release was 100 %. We examine a snapshot taken 5 years after tracer
release. </p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f05.pdf"/>

        </fig>

      <p>First, we examine four simulations where the DP remains <italic>closed</italic>.
Perhaps as might be expected, unable to cross the TG and lacking alternative
routes, tracer “species” remain on the side of the TG where they were
released when the TG remains closed (Fig. <xref ref-type="fig" rid="Ch1.F5"/>a, b).
However, when the TG is open, tracer species released from the
<italic>west</italic> of Australia (Fig. <xref ref-type="fig" rid="Ch1.F5"/>c) enter the AAG
along the Australian coast but do not proceed into the Ross Sea, east of the
open TG. Instead, concentrations are focussed along the Antarctic coast in
the South Atlantic (and west of the AAG). This is due to the westward flow
along Antarctica there and in the AAG (Figs. <xref ref-type="fig" rid="Ch1.F1"/> and <xref ref-type="fig" rid="Ch1.F4"/>). Westward propagation along the Antarctic coast of
dinoflagellate tracer released <italic>east</italic> of the TG in the open-TG
configuration is evidenced by the local maximum in tracer concentration along
the Antarctic coast of the AAG in Fig. <xref ref-type="fig" rid="Ch1.F5"/>d. This is
also because of the westward flow through the TG (Fig. <xref ref-type="fig" rid="Ch1.F4"/>). The
biogeography that can be inferred from our results is in agreement with
<xref ref-type="bibr" rid="bib1.bibx6" id="text.37"/>, who infer that lower-latitude taxa that are abundant to the
west of Australia remain geographically constrained to the west of the TG
after the southerly opening of the Tasman Gateway. Notably the consistency in
the simulated biogeographic patterns inferred from the model is unrelated to
the regional environmental differences (e.g. temperature) and only the
result of current vectors distributing the dinocysts in the region. This
absence of a relationship between environmental factors and biogeography is
particularly convenient since <xref ref-type="bibr" rid="bib1.bibx5" id="text.38"/> also showed that the onset of
regional dominance of endemic dinocysts in the SW Pacific Ocean was unrelated
to surface water temperature. Also in agreement with field evidence
<xref ref-type="bibr" rid="bib1.bibx6" id="paren.39"/> is the westward propagation of SW Pacific tracer species into the AAG in response to gateway opening and the restriction
of that propagation to the Antarctic coast of the AAG.</p>
      <p>Second, to elucidate the role of the DP in our dinoflagellate tracer results,
we performed versions of the second set of tracer simulations
described above where the DP is now consistently <italic>open</italic>
(Fig. <xref ref-type="fig" rid="Ch1.F6"/>). Similarly to the DP-closed scenario and
perhaps trivially, when the DP is open and the TG remains closed, species
released at both sides of the TG land bridge remain on their corresponding
side of that land bridge (Fig. <xref ref-type="fig" rid="Ch1.F6"/>a, b), although very
small concentrations penetrate north of Australia via flow through the open
DP. When the TG is open, tracer released to the west of Australia
(Fig. <xref ref-type="fig" rid="Ch1.F6"/>c) enters the AAG. However, unlike the DP-closed case, tracer now penetrates well into the Ross Sea. This is because of
the eastward flow through the open TG that is part of a proto-ACC flowing
through the open DP (Figs. <xref ref-type="fig" rid="Ch1.F2"/> and <xref ref-type="fig" rid="Ch1.F4"/>). This
effect is in strong disagreement with field evidence, as dinocyst assemblages
in the SW Pacific only show a low-latitude affinity when the Tasmanian
Gateway deepens at 35.5 Ma and not earlier.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6"><caption><p>As Fig. <xref ref-type="fig" rid="Ch1.F5"/>, but for the Drake Passage open. When
the Tasman Gateway (TG) is open, dinoflagellates released to the west of
Australia enter the Australo-Antarctic Gulf and flow into the Ross Sea
through the TG <bold>(c)</bold>. Species released near the Ross Sea may enter the
gulf along the Antarctic coast <bold>(d)</bold>. We examine a snapshot taken 5
years after tracer release.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f06.pdf"/>

        </fig>

      <p>Some westward propagation along the Antarctic coast of dinoflagellate tracer
released east of the TG in the open-TG configuration is evidenced by the low
but discernible tracer concentrations along the Antarctic coast of the AAG
(Fig. <xref ref-type="fig" rid="Ch1.F6"/>d, compare to Fig. <xref ref-type="fig" rid="Ch1.F5"/>d:
note the smaller scale in the latter), while most of the tracer flows north,
along the east coast of Australia, in the Ross Sea gyre. In conclusion, the
tracer dispersion patterns in the DP-open case are not in agreement with
<xref ref-type="bibr" rid="bib1.bibx6" id="text.40"/>, as significant amounts of tracer released west of Australia
penetrate into the SW Pacific upon the opening of the TG. This indicates
that either the DP was closed during this period or sufficient obstructions
existed downstream of the DP to prevent an eastward flow through the TG.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F7"><caption><p>Sea surface temperature change in response to opening the Tasman
Gateway (TG). Difference between TG open and TG closed for <bold>(a)</bold> the Drake Passage
(DP) closed and <bold>(b)</bold> the DP open during the opening of the TG. The
opening of the TG leads to a maximum of around 4.0 <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C localized
surface ocean cooling to the east of Australia only when
the DP is open.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f07.pdf"/>

        </fig>

</sec>
<sec id="Ch1.S3.SSx2" specific-use="unnumbered">
  <title>Temperature changes in response to gateway opening.</title>
      <p>The opening of the TG in our model configuration where the DP remains closed
leads to no significant sea surface temperature cooling (Fig. <xref ref-type="fig" rid="Ch1.F7"/>).
This is in contrast to the hypothesis of <xref ref-type="bibr" rid="bib1.bibx6" id="text.41"/> that the cooling
seen in their field evidence of surface water and regional air temperature
reconstructions is a direct result of gateway changes and the ensuing changes
in ocean currents. Furthermore, none of our experiments show significant
deep- or mid-ocean cooling. We suggest that future climate modelling work could shed
further light on the relationship between Antarctic temperature changes and
the gateway changes. Sea surface temperatures close to Antarctica have
similar values of 10 to 12 <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C on both sides of the TG when the TG is
closed and the DP is open or closed (Fig. <xref ref-type="fig" rid="Ch1.F8"/>). As a result, although a
westward flow emerges from the Ross Sea to the AAG (see above), the opening
of the TG in our model configuration where the DP remains closed leads to no
significant sea surface temperature cooling (Fig. <xref ref-type="fig" rid="Ch1.F7"/>).</p>
</sec>
</sec>
<sec id="Ch1.S4" sec-type="conclusions">
  <title>Summary and conclusions</title>
      <p>Our model results provide a numerical underpinning of the recent field
observations and interpretations of <xref ref-type="bibr" rid="bib1.bibx6" id="text.42"/> that a southerly opening
of the Tasman Gateway causes throughflow of SW Pacific surface waters into
the AAG. For the first time, we reproduce a westward propagation of oceanic
properties and species originating east of the Tasman Gateway upon its early
opening, leading to the introduction of endemic dinocyst species originating
in the SW Pacific into the southern margin of the AAG but not the northern
margin. This is consistent with our finding that the westward current is
restricted to the southern margin of the AAG. Also consistent with
micropalaeontological observations, no low-latitude-derived species (released
west of Australia in our simulations) could be routed via the TG to appear in
the SW Pacific Ocean upon its opening.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F8"><caption><p>Sea surface temperature for <bold>(a)</bold> the Drake Passage (DP) closed
and <bold>(b)</bold> the DP open.</p></caption>
        <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f08.pdf"/>

      </fig>

      <p>Importantly, our model results indicate that the waters of the DP, or
upstream or downstream areas close to it, are likely to have been obstructed by large-scale geostrophic flow during the early Eocene, as the passive
tracer experiments in a scenario with a closed DP are much more consistent
with microfossil evidence compared to a scenario with an open DP.
<xref ref-type="bibr" rid="bib1.bibx25" id="text.43"/>, <xref ref-type="bibr" rid="bib1.bibx17" id="text.44"/>, <xref ref-type="bibr" rid="bib1.bibx13" id="text.45"/>, <xref ref-type="bibr" rid="bib1.bibx27" id="text.46"/>,
<xref ref-type="bibr" rid="bib1.bibx14" id="text.47"/>, and <xref ref-type="bibr" rid="bib1.bibx28" id="text.48"/> infer a progressive opening of the Drake Passage,
through continental extensional tectonics and oceanic spreading
<xref ref-type="bibr" rid="bib1.bibx15" id="paren.49"/>, after about 50 Ma from the analysis of seafloor magnetic
anomalies in the Scotia Sea and adjoining oceanic areas. This timing suggests
that the DP may have been sufficiently obstructed to prevent an ACC.</p>
      <p>Our results, and those of <xref ref-type="bibr" rid="bib1.bibx6" id="paren.50"/>, indicate a later opening of the
DP. Alternative to the DP closure, we raise the question (for future research)
of whether a similar effect could have been achieved from severe obstructions to
a wide and deep flow through the DP and nearby areas at similar latitudes, a
possibility not explored in our model experiments. For instance, model
results by <xref ref-type="bibr" rid="bib1.bibx19" id="text.51"/> indicate that, regardless of the state of the DP
and TG, a coherent ACC was not possible during, for instance, the Oligocene (a
period much later than that under study here) due to the Australasian
palaeogeography, although no inferences are made for the Eocene. Future work
on these obstructions is therefore important. Finally, our numerical study is
not consistent with the idea that such an oceanographic change can cause a
significant and uniform Antarctic cooling. Nonetheless, our result of a
westward current made possible by a restriction of the ACC away from the TG
region provides an ocean dynamics background to the finding of
<xref ref-type="bibr" rid="bib1.bibx6" id="text.52"/> and other studies that point to this current.</p><?xmltex \hack{\clearpage}?>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <title>Dependence on model and geometry</title>
      <p>As the position of Australia plays an important role in whether the
subtropical and subpolar gyres of the Indian and Pacific oceans pass that
continent northward or southward, we performed additional simulations with
(i) another ocean model and (ii) a different reconstruction of the
continental geometry. We use the Parallel Ocean Program (POP) developed at
Los Alamos Laboratory <xref ref-type="bibr" rid="bib1.bibx12" id="normal.53"/> at a nominal horizontal
resolution of 1<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> <inline-formula><mml:math display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 1<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> and 40 vertical levels. The
model was adapted to a late Eocene reconstruction of the continental geometry
and bathymetry <xref ref-type="bibr" rid="bib1.bibx34" id="text.54"/> and forced with the atmospheric state of
a coupled climate model simulation using the Community Earth System Model
(CESM; <xref ref-type="bibr" rid="bib1.bibx18" id="altparen.55"/>). Compared to the continental geometry in the UVic
simulations of the main paper, Australia extends substantially further north
in this configuration as it represents the late Eocene or early Oligocene
rather than the middle Eocene. The model is run for 570 years in two
configurations: the late Eocene control case, where the Tasman gateway is
open and the Drake Passage is almost closed (35 m deep) and a second case where
a land bridge is built in the Tasman gateway from Tasmania to Antarctica. The
throughflow through the Drake Passage is very small, so we can consider the DP closed
in these simulations. We show results averaged over the last 10 years of
simulation.</p>
      <p><?xmltex \hack{\newpage}?>Figure <xref ref-type="fig" rid="App1.Ch1.F1"/> shows
the vertically averaged zonal velocity for the two cases, with contours of
sea surface height (SSH) overlaid. The SSH can be regarded as similar to the
barotropic stream function as it represents the barotropic flow. In general,
the POP simulations show very similar features to the (coarser)
UVic simulations shown in the main part of the paper. The model is forced by
the same wind pattern as the UVic simulations, and in particular, the Tasman
Gateway is entirely in the latitudes of the polar easterlies. However, in the
POP simulations, Australia is located further north and, therefore,
effectively blocking the eastward flow at the boundary between the
subtropical and subpolar gyres. As a result, less flow passes north of
Australia from the Indian to the Pacific Ocean, but a substantial part of
this flow also passes south of Australia, thereby inducing considerable
eastward flow in the Tasman gateway when open (Fig. <xref ref-type="fig" rid="App1.Ch1.F1"/>b). In a
vertical section through the Tasman gateway (Fig. <xref ref-type="fig" rid="App1.Ch1.F2"/>), this becomes evident as a strong and deep-reaching
eastward current in the central northern part of the gateway, while a weak
westward current at the Antarctic margin remains. This gives further support
to the notion that, if the DP is sufficiently obstructed, the eastward flow
at the boundary between the subtropical and subpolar gyres returns mostly in
the subpolar gyres, i.e. along the Antarctic margin.</p>

      <?xmltex \floatpos{h!}?><fig id="App1.Ch1.F1" specific-use="star"><caption><p>Annual and vertical average zonal velocity (cm s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>; positive
values indicate eastward flow) with contours of sea surface height (SSH in
cm) overlaid for the late Eocene simulation using a different ocean model
(POP), with the Tasman Gateway (TG) closed <bold>(a)</bold> and open <bold>(b)</bold>.
The Drake Passage is shallower than 35 m in these simulations and can be
considered closed. </p></caption>
        <?xmltex \igopts{width=369.885827pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f09.png"/>

      </fig>

      <?xmltex \floatpos{h!}?><fig id="App1.Ch1.F2"><caption><p>Meridional section of zonal velocity <inline-formula><mml:math display="inline"><mml:mi>u</mml:mi></mml:math></inline-formula> (cm s<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) inside the
Tasman Gateway for the late Eocene POP simulation. </p></caption>
        <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://cp.copernicus.org/articles/12/807/2016/cp-12-807-2016-f10.png"/>

      </fig>

<?xmltex \hack{\clearpage}?>
</app>
  </app-group><ack><title>Acknowledgements</title><p>We thank the University of Victoria staff for support in usage of the their
coupled climate model. This research was supported by the Australian Research
Council (ARC). This work contributes to ARC project DP1096144 held by Willem P. Sijp and
ALW 802.01.024 of the Netherlands Organization for Scientific Research (NWO),
Earth and Life Sciences held by Anna S. von der Heydt. The POP computations were done on the
Cartesius computer at SURFsara in Amsterdam. The use of the SURFsara
computing facilities was sponsored by NWO under the project SH-209-14. Peter K. Bijl
acknowledges NWO-ALW for VENI grant number 863.13.002. Figures were produced
with the Spacegrids Python data analysis module. <?xmltex \hack{\newline}?><?xmltex \hack{\newline}?> Edited by: A. Paul</p></ack><ref-list>
    <title>References</title>

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    <!--<article-title-html>Model simulations of early westward flow across the Tasman Gateway during the early Eocene</article-title-html>
<abstract-html><p class="p">The timing and role in ocean circulation and climate of the
opening of Southern Ocean gateways is as yet elusive. Recent
micropalaeontological studies suggest the onset of westward throughflow of
surface waters from the SW Pacific into the Australo-Antarctic Gulf through a
southern shallow opening of the Tasman Gateway from 49–50 Ma onwards, a
direction that is counter to the present-day eastward-flowing Antarctic
Circumpolar Current. Here, we present the first model results specific to the
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shallow opening of the Tasman Gateway indeed causes a westward flow across
the Tasman Gateway. As a result, modelled estimates of dinoflagellate
biogeography are in agreement with the recent findings. Crucially, in this
situation where Australia is still situated far south and almost attached to
Antarctica, the Drake Passage must be sufficiently restricted to allow the
prevailing easterly wind pattern to set up this southerly restricted westward
flow. In contrast, an open Drake Passage, up to 517 m deep, leads to an
eastward flow, even when the Tasman Gateway and the Australo-Antarctic gulf
are entirely contained within the latitudes of easterly wind.</p></abstract-html>
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